IN THE NAME OF ALLAH THE COMPASSIONATE, THE MERCIFUL

IN THE NAME OF ALLAH 

THE COMPASSIONATE, THE MERCIFUL

TAXONOMIC STIJDIES OF GRYLLOIDEA 

(ORTIIOPTERA) OF PAKISTAN 

.RY 

MUEAMMAD SAEED 

M.k.(HoN.) Asn. E MIog, 

A th.is tuhnitt.d k Naiat futlbn ',t ol 

hc rcquircn0ts Jol th..ktra of 

DOCTOR OF PHILOSOPHY 

IN 

, j terucwtaut ENToMoLocY 

FACULTY OF AGRICULTURE 

UNIVERSITY OF AGRICULTURE 

FAISALABAD 

IPAKISTAN) 

'1995

To 

Th. Codollcr of EirDidiorE, 

Unilrnity of ASrlorltur., 

wc. rhc SupcNisy Cn|nhin e, crdry $$ 6r dlllnt &d imm of 0Erit 

lub|tried by Mr. MdlrtnrEd S..d haE bccn fould ! i!6dory.d ntomrEnd tlul 

n bc prs.d to cvrlurrh by dE En.'nd E'diF(r) fo. tu $ntd of 0t d.g!.. 

CHAIRMAN : 

MEMBER 

MEMBER 

SIIDERVISORY COMMIITEF. 

tIgEABy: 

MUHAMMAD YOUSUF 

DR. SHAMSTIAD AI(BAR 

IUNAID IQBAL QU

DEDICATED 

To Dy hc poftol!, 

rnay h.ir rorb rE3t ir p.ace ir H€lvctr!.

Dar Mr, M!h!a.d Arr&S 

colnollcr ol Brrnin$@, 

I r!ii., rb tt.D. dEi. orMt. Mtud sd b dffid.. El brd & rd o, 

Ii.D. d.arE, 

I Itt@ tb tbL, d. dd-.btl bc rt ..dd.d tu r! agrl aftb&aF. 

t!.rd!.q b b . F!@ wno b. $. thod t!. doidio (flEd)

Der Mr. Muh.nnad Alhf.q 

Cmmlhr ol &mindid, 

Enernrl E (rtrlln€r'r Commcnts 

| [* @tutlr E:d rb. lb6L of Mr. M. S...d .d gi* oy npdt h.le, 

I fild, ric dai.6lly tultlls rll ti. Eqrl!@d fa I tn.D Lwl lgd t qli|ltld I! 

tul, | @y ey tt:r I fi'n it d oiLidii'8 pke or wL 

Mr, s..cd hs d.@tnr.d his @nDkE uodcsludiry dtlF Dlhodt of l,Io|mic @@h 

ge hs .xoircd au or in ELvrr liErdoud hs nis!€d @tlln8 H. ba rhM bj! 

twlcdgG ol $c imporl$t cts&tets us.d ln dE l"rdmt of crickai. Thi! ir o inponam 

srdD of leti rol olrt b€.w ttcy cru& @Bijcrabl. dxlmic &D.a! t0 cr0p3 dd 6torql 

product!, u eEtully poinr.d ol by Mr, $!.d, bu1.l5o bqeE tb.y c ! ioponrn sop 

doluliMily. W. e jBt b.siminS to F tw l4p atd divd l[i! &op or it!.ca b. As 

...d by Mr, S...{t, l6n .ndi.5 in A|6li. .nd oina rad p.r!. oa tL wld tut36a the 

6lla! e I v.ry l{9. am!. Th. rumba ofw rFis {b!dib.d by Mt sd tn6lto. 

qaidcnbh f.@. ir P*isro. 

Mr. SGd hs dcmltratld thr! lE Do$.{3.3 . Ey ad.pr td!nq!. iI ttudylnS cri:t rs Tn 

e of m.L i.nirrlir iD dtumy ot cricL! b . Elrliwly @lt .djutr t to tndy, Mr. Sae.dt 

dnwin!! shor l|.|. inponet ch{a.t s much bai.r lho 3imila lllu![.doB of @v noE 

I cdlntul.l. Mr. Sr.cd @ . beltitul !t!tl, shnn I wh h. hr!'y to h.s dN Dt*tf r 

*qld liL io .|Mr4! nio !o pttD$ n fo. Pbli.ri@ dd hA. rha t wlll h. .bk b 

snyu3 d. odaio of P:rllr. 

"sd-" 

I n w d.d. r Gy &w @4ioal c.tMn i! dr body of tt thdit rye|t tn6. in @ sy 

.Lr.d nm |a inpon !@. Ako p.!c 84 w nisiIg fror nt cAy. fiow6, I u 6oi! it 

wlll b. of rlE se qualitt 6 thc orb.r DUq .!d do lot tr .d to !..d lr i, ord.r ro ps

ACKNOWLEDGEMENTS 

I h.v. no *ord! to cxPBs my dccFst scffc ofgl?tiudc to Alnthtv 

Alhn mhc Mcrcittl and Grdcious), who cnabl€d Inc to complctc thcs€ sodics- 

I lrhh to cxPrcss my thttb ro my luFrviso. Dr' MunantEd 

Yousuf, Pmfessor of A8ricultunl ErtornoloSy, University of ASricular.. 

Faisalabad, for hh subslanhal 8uidancc, continuous inspiration al'd invalurble 

assillNrrr in bringing lhii diss.rlalro. to il3 presltrt fo.m- I am Stcdly ind€b|d 

to lhc mdbers of my $p€ohory comiu.c, Dr. Shamshtd Akbrr. P.ofNr of 

Agriculfural En omoloSy and D.- JMid Iqbal Qureshi, Professor of zoology, for 

dlcir valuablc su88€slio$ nd coBtucliv. criticism throughoui ihis wo.k 

My since.e thrntr are dE to Dr. Dmicl Olie. Acadcmy of Natunl 

S.i6q of PNlad.lphia, U.S.A., Dr. A.V. Corothov, Z@logicrl ltsdaitc, 

r-.ringrad, U.S.S. R. , Dr. R.B. Toru, Tmvaal Musam, Pretori|, South Arrica, 

D.. Yduhiro Orhirc, M&jima, Naha, Japan ad Mr. Khalid Manmood, Ph.D. 

slldmt at thc Na0rrd History Mueum, lrodon, for Ileir kccn int€.61 and 

Scncrous hclp in s€ndin8 thc valuable lilcranlre on gryllids drinS this work. I atn 

ako ll)anftnrl to M.- Azlul Saed, l€t!ftr, D.paimEn( of AS.icultunl 

Entomolosy, UniveBity of Agriculturc, Faisalabad, for 8ivi.8 his id..tificd 

sDccim€ns fo. comDartuon and litemtrrrc ofl loan.

t ftd SrcNt pl..ttlt€ to c cflt my lppftriitiont !o Dr' L D$utt'r- 

Grrndcolat, Mol6|n Ndrtl d'Hlsioirc Nrnltclc' Paru snd Dr' D C F Rd ?' 

cslRo, Divtuion of Enlomolo8y, Cj!'bcn, Au'!']i$ fd dEir crittc'l collund s 

ad kxtnical sdvicc to improvc thi! drflrlcript' 

t tm cap€ciilly tha*fuI to ny rlc5|ch collca8uc'' Mr' Muht nad 

lqb.l !!d M. Hui|lin Z!fir' wtr 0!&Gd oc wilb tid' td d€rrv dt'itg |[e 

freld co{.dhru. My rppr.ciirion tu slto dlc to tnv friqd ' Mr' Alif S'ccd' for 

ryping thi! lttolcripa. 

FiorIY, tE 

dhor crprt$.. hir d.cpd gdilt& @ bb hnilv 

m.obc$ for dEn morll |Dd fiD&cirl JjDfoit thrflS]flt ni! rdcmi(: ctrccr'

CONTENTS 

CIIAPTER TITLE PAGE 

INTRODUCTION .......... I 

II REVIEW OF LMRATURE 

III MATEruATS AND METHODS 

RESULTS AND DTSCUSSION ...... _....,..,....... t3 

SUPERFAMILY GRYLLOIDEA.............. 13 

Family Gryllidae ........................ t5 

Subfamily Cryllirue.... ............-t7 

AeN Ttbii*i.lha ................20 

AcaB PhoM.Ila . ............ -....22 

G.not Tunlog,itts ................25 

Gcna CallbSryltu .................43 

C.tis ArUopsis . . . . . . . . . . . . . . . . . . . 46 

G.M O/rl&J . . - . . . . . . . , . . , , . . - - .47 

cFw T.l@Errltu ...-.............50 

Gav PLttrytta............. -....6E 

CEM PhallolobM . . . . . . . . . . . . . . - - 76 

C.nw A.heta.. -..................83 

C.na PLb.iott Uu ................91 

Cenlt Ctraodct . . . . . . . . . . . . . . . . . . . 94 

G.nB Tat @tryut t . . . . . . . . . . . . . . . . 96 

AeNs 16ob1M................ 102 

G.nt v.loifcrorc... -. -.......... t46 

G.N Modi@tryUB .. -............ l6E 

Subfamily Nemobiina€................ tEE 

A.nns Pt.tuh.n bilt ,,.........,... 189 

A.na Diaartubi6 .,...,. -.....,.. l9l

CHAPITR TITI,E PACE 

Fnnily Trigodidiidae ...... . ... ... .... .. - 195 

GeM Anlltgus ................. l9? 

ctNt TnSo!,Au - . . . , . . . . . . . . . - . 201 

CrtN Awipha .................. 206 

Frtrfly Phalangopsidae . . . . . . . . . . . . . . . . . . - 20E 

ca6 M.loituah4 , , . . , , . , . . . , . . . . 208 

Family Eneopt€ridae ........ -........... 216 

Subfemily En€opterinae . . . . . . . . . . . . . . . 216 

CnNt Xdotryllu , . . , , . . , . . . . . , . . . 216 

Subfamily Podoscininae. ....,......... 2r9 

G.flie &.rdrr6 . , , . . , , . , . . - . , . , . . 219 

Famify Oeca hidae .....................22r 

ceus obamtB . . . . . . . . . . . . . . . . . .221 

Family Mogoplisddae . . . . . . . . . . . . . . . . . . . 233 

G.dB Gott'dla ................. 233 

Farnily Scle.opferidaa . . . . . . . . . . . . . . . . . . . 23E 

Cen'rs Sclztopt M ......,,........ 238 

Family Gryllo@lpidee ................... 2fi 

Gents CttUotaba . . . . . . . . . . . . . . . . . 240 

SUMMARY . , , . . , , , . . , , . . , , . , , . . , , . , . . . , , . , , 250 

VI CHEC(LIST OF CRYLIOIDE,{ 

(ORTHOPTEM) OF PAKISTAN 252 

v LITERATURE ClrED ,. _ . , . . . . . . , . . . . . . , , , . , . . .56

L 

28, 

3. 

5. 

6, 

1, 

E. 

LIST OT'MAPS 

PAGE 

Con.cio6 lrcalnk of Crylloidc. of Ptisrn. .............. 9 

DiEibltion of Grylli'|,a lPhoMnur.t, IBrrrogtUat, 

CaUoeryllw, CryWh, eryUllt, f.hoStrAB, Plat styutt, 

Phalolobow, ,rcha. aM Arylwrs) In Patirtan. .......... 255 

Di'fiblulid of crylllnt lTtutuogrylrd, laobhw, 

v.larif4t4u d MdleoSrylw) in P*i!nn. ............ 266 

Dirrributid of N.nobiin* in Pti'trn. . . . . . . . . . . . . . . . . 267 

Dtnbu*n of T.igonidiidr in P.t!lrn. ............... 268 

DlEibutb of f,bhngop.i.n, Sdsqrdil& ||d 

Mogopliiirj& in P*bon. ........ _.. _..,.......,. 269 

Dilr.ibudon or En opErftlt. in P*h!l' . ... ... .. .... ... 2m 

Disr.ibutlh of oo.dihid& in P*i!l!!. . , . . , . . , . . . . . . . . 2?l 

Dbt ibutio ofcrylloirlpnt. in P!kir!n. ............... 272

LIST OF FIGI,]RES 

FIGURE TITLE PAGE 

9. 

t0. 

ll. 

t2. 

13. 

15. 

17. 

18. 

19. 

20. 

2t. 

Prircipal trupholo8idl fcrn A 

G.ylloiiia (Diff@nt fanil'ar). 

cryilida. (Diff.renr subflmrli.r). . . 

of rh. dicllcr. ........ .... 12 

PhoMftuB ni@t tF?dt, .lym atd gcniblia). . . . . . 

Tr tu try I lB s a Unat6 (EtYa\ 

fwst 1116 pakist@N lH.d. pronoton md clyta) . 

Ttt@tryU!.t kolawLtit sp- w. (F!... hed, prcnoilh 

rt@eDU!4 b.!lah' .rp. mv. (F@. h€5d. Prorcom. 

eryua and ovrpGrtor uNat. 

Iu@erytlls bdtali 3p. w. {G.nnali{). . . . . . . . . . . . 

Tu@eryU!6 pndi.B sp. w. (Fe and tlyt!-a). . . . . . . 

CdlloerJl8 ovtuaeB !p. B. (F@, hqd. prono(m. 

.lt,ra. ercB lnd oip6ilor). 

e'tLbt biwlatw (Giialia). ....... 

fcl2oery la USM6 ep. n@. (Eltr. and geni6li.). 

r.teotryll6 tonSiptwns lEtyra ud gcni6ti^t. 

r.t osrruu toB6 !p. w. (Elrr'r.rd 8cni|.li.). . . . . . . . . . 

T.koeryllla ptu rB 3p. rcv. (Elyh &d gFni6iia), , . . , , . . 

T.l@eryll8 laaeuid$ sp. nov. (Elyn md gcnitrlia) .,...... 

PIary{rUB bMn (Gcn|hlb ard qipcilor valB). ....... 

PI4Ntt lttg ulqnpta .p. mv. (Fa... elym, ov'pditor 

larvB ao tcnMrE). 

PhaUobb$B ad.ntibiaut 8en. a 3p. rcv. (Elyb. aJd 

sohdwis Een. d sp. mv. (Elyln .rd 

2A 

2E 

30 

33 

16 

36 

45 

49 

56 

59 

63 

67 

70 

8t

FIGURE 

22. 

27. 

25. 

26. 

27. 

28. 

10. 

ll. 

17. 

TIII,E 

(Head, Prcrctum. 

dYtb and 

sp. @. (H@d, Prorctum. f@ clt@ 

TanarosryIlut tunarLt lFM, head, pronotur, .rv(la 

29. Isobt "n!!t .rcabpta sP nov- (Hcad' prcnotun' 'lviB 

12. Ldobknnlt nuai6.phot)B + iot tqt?d.qrcnoun 

he, Itt lnt nnrl sqmlnt,.lvh and cprPh.llu,. 

34. 

35. 

36. 

38_ 

Pkb.ioSryll!4. Hn gulatit sP 

.ryhaJrd gcrrr.rd). ...... 

Gtylttd$ !Wu.a6 tctnibti^) . 

and g€niolia). . - . , . , . . . - 

IEobIwB potohse8is sp. nor' 

cryn). ..,.-..-.,.,..- 

Idoblqtld doli.hory4nis sP. 

Lotobt.MB oneubs6 sp. nov. (Elyut and S.nitali!) . 

Idobl.nn's .o46a sp. d. {Elyr' ad 8.ni6lia) 

tdo ctturu.s .ktw,,ato sp. N. (Hod. Prcrctom, clvh 

andSeni6lia). ......... 

broblatu til*en6 sp. nov (Hqd, prcnotum. l$ 

anann l sgmc , .lyh lnd g.ni6li.). . . 

Lotobkw h.NhMLB 4, mv. ( lst antcndl *8m.N 

LdobkMq haruuf.t ..(tlezd. pronotum. lst lnl.nMl 

i.8denr, clyFa and g..irdiz 

19. vclo.ifictot s atveotB sp. mv. (H.ad, pronotun, clyr. 

andsenii.lia), ........ 

40. vetatifutorb 6pa^d lHqd. promtum aJd g.niblia) 

PAGE 

85 

8E 

93 

91 

100 

t0l 

r05 

t0E 

l12 

lt8 

.. t25 

..110 

t.ll 

tJ1 

t4t 

152

FIGT]RE TITII PAGE 

42, 

46, 

50. 

51. 

52. 

54, 

55. 

56. 

51. 

5E. 

veloifi.ro4 Unnn66 6p. nov (Head, prcnotun clvtm 

and cpiphallut. ... ...... 

V.brif.rotu tztittut8 tp. nd, tHad, p.onorum, .lylrr 

43. v.loiftcnw cotun B !P. w (H.ed' prcmun' .lvda 

45. 

59. 

60. 

and.piph,lls). .......... 

v.taif.to$ lall4t lE+iptatt'!s). 

a. mv. (H.!d, Prcnonm, .lyt . ud 

Benihlia). ............. 

Modi.ogrya^ bl.nnu (Et!tt^ ^rA 8.nid,^). 

Motli.o8rytk( tit\tlto8. (Elyrre.d geniblh) 

Modi@gty !6 .l$ar (elyo and Scnialia). 

Modicoeryllls bo@rLtt sP. nov. (F@, elvtt.a and 

MdktSttl'la ..r.t l4 e. @. (F&. clyra ed 

Mo.dicgqrtaq ry19&P9n!. 

ovrposltor d6 M 8.nBrsr. 

sp. d. (F&, clYr.r. 

Dia4.refi6 didi (Elyra, P6terior femu &d post4id 

ribia). ................ 

A46u4"6 *trrut$ sp. rcv. (H€ad. pronoom. clv@ 

oviposns dd genilali!), . . . 

Tliqot4dittn cicird4gld.t (Elytr4 and g.niadi.), 

r. c. hunbeniM \El,JlJt). 

Awipto lon$ip@it (EtrE^). 

M.loiM'pha ne oôta tp..tov. (Hed, premtun, clyr.. 

.rd g.nitalia). .... .. 

E6.ynu corciM6 (Po.btiol ribia). 

Oeeanthlt irdi$ (Sfidrtatory 

@.(Elrra)...-............ 

(Elrlra, p6le iq libia and 

154 

t58 

t62 

164 

t67 

lr0 

t13 

175 

t79 

ta2 

186 

!93 

2m 

202 

201 

2tt 

2t1 

2t1 

no 

224

IIGI,RE ITI'I,B PAGE 

61. 

62. 

63. 

65. 

6. 

68. 

69. 

Oeot hB cttut\l4t |o, N. (Gl||iuLt pit of tllc 

Ilr.rb$n. clyr.r id saidll.to.y nh). . . . . . . .. . .. 

tud'dt"s cnmt Lt |P. rF,,. (G.nirllr) 

O@tt,/,t datwmnat s. ftr. G 

ncr.ndun, clrr. ind rkhilttcy lilo). 

O..afltutt d.tu p,,ttut ry. r!v. (Ctttirli) 

Gar'radt otadi,natle 

prc!m||n, olyrh .!d $dotun). 

Q. |Dv. (Fc, E d. 

$l],,ptctt-c. pffi,'It (rbd, p(odrE, f&, dtut.td 

paE !. nDrr. 

G,yll,el,a oda@b (Blyfr .d i.nulm, fllc). . . . . . . 

C,fllelry ,rl',,,' (El'ln.d Lld'ttD.y filc). 

Gtruotob. t&M rD. |!v. (Pr!|lotm. clvt t, 

Doiic.kr'lcmui, tl'n"b..y |ilc rn $midr). .. ......... 

226 

tn 

29 

232 

235 

2t9 

u2 

24 

u7

INTRODUCTION

CHAPTER I 

INTRODUCTION 

Tlc mchb.rs of thc auperfanilv Grvlloidca se onnonlv 6llcd crickcts ehich 

de qidcly dhtribuad. mGr abu.dant and with . wide |ange ot food Plrna ln 8crcrd' 

$ey ate coBidcrcd b be odnircrcls iisc.s 

Orylloi(l3 have s@t @mnic impo !@. Sone @ impona p6ts of fEld 

crop!, veSdabLs, l.wN, (llrjaftnlal pla s. hfsi.d gdiN bo{h at thBhing flla ard 

godowN, and hou!.-hold dicles in diffcrett Puts 

of $e wodd For erdPlc, $' adul$ 

and nlnphs of !hr@ rprie!, vn Ach44 .lotulk4 AdUB bi@ulatt ^nd 

Grhno|rylfut erlthrteph4Lt^, plav havo. bv fceding voracioulv on e'd aDd s'dlings 

ol couon. mill.$ and oil-s€€

4 serious peea of pornroi iob@, ri@, r.r {|d abur garden pbnB in o. IndcP.t 

3ub@nircnt (Cho6h, l9l2i Chopdd, l96q Anw.r dL, l9aD. S.w6l sp@i.. of rh. 

8.nE @lllotalpa harc ben ilport d d.masiB v..iou! ctopr, hs, i..d b.d! .rn 

omamenlll planb (Iown6cnd, l%3). C. lNol, for cenple, i! a srids rcst of 

pobto crop in W6t 8.r8d, Iidia (B.Mir at|d Cn|d.te, 1955r, whi|c G. 4ri@ 

d.,rag4 mB or Chine. Yam in .r.p.i (M.tsuuh .r rL, 1985). Even rhe clrnivoos 

sp4id d eue exteNivc mdh{iel &nag. to cogr by thct burNing &dvhy 

(M!lfu'y, l9E l), wh'ch alo 6drs in &n ge ro $c bants or w.r.r chatrls .!d dcylc 

(rindale. r92E). 

Crick.6 e alio p€s$ of he6r.d gaiB ard c!@ hany 169 by @1ing rh.n 

on the rhBhing floors. For.xantqt , Ct l,la donatiN prcf.B to f..d on rhc t6iB 

of nal'rd, what and srshum alongwhh $e semiMting khdif crops in LrkaM, 

Drdu, &.nhi (Kalar sial!).rn UsrN Muhanrud (Brluchiran) in PrtiJnn, Ir mrrully 

6t! aMy 0.5-2.0 Fr cent of wh.ar ghiB ud 6 Fl € of oil{ed3 pe. nighl €uing 

a. ov.nll ld of lG25 pd @.t pcr nonth on the thBhirg nl$ (AnDd and 

Ull.tl' l9l9). 

In houFs and sodwns, e'JUorles isillatu ^tt A.hda done$ica caw 

@Bidehu. dMlge to lh. 3lorcd f@d! !d orhcr mtcriah lilc whar flour, bi.rd, 

Soo.iim*, thc dick.tt navc allo plwd vcry @ftil du. io ihcn f..ding on 

impdra irs.d pesa. Fd mplc. G binwahn t hx bc.n s€.n fc.di$ on 

grahopp.B atd catrpiuals (Amn., Itl-73), shiL G. donEsthut ad Ary d6 

tSillard f..don!henipt ls siored Snin pq! Aphand 4itto?l,lh, atd\E.Ege. A 

3ingle qiclct could dr a 'fuy d 25 cge! in ld rlEn ev.n hom (Cdbr, l9t6). 

Siniltly, Cryllotalpo E tllotalpa ha! somctinA b.eo nocd f.€ding oa 3mall itMa dd 

wom3 in diffcent partr of rhe @rld.

-3- 

Th. Eromrny of ti6. iDpod.nt lt!..t' hs bcn craEiEly crDldld ln &. 

Orintal Esioo includnE lldit by . nunt r of Qs*.6, bd io .vttematrc |lan96 

p.uining ro th€ grylbid ftura of Prtbul hrvc bcen md. .xc.pt thd. of Chopdrd 

(1969), Abdullah 0955) rflt S.4d (1969. Tli. fd@ id.nrift d odv la lpeict out or 

r r.6l of 355 ryei.! frlo P.rh.d, ehib ttE l.!a rs did ff.t of Err PcliEintv 

lnd nSionl ffirc l|l|t t o icded ftlan t codrnlibG. Morlovq' t ddsifE dor 

of this group hs und.rSoE 6!@ttdou! chDg.! rt.ulting in thumiry.td BhuminS 

of v.riou raxa in dl! rcclnt Posl 

In vis of tlit, d dEsitt tlrrcy w di.d ol iom 1991 !o 195 to 

.rploE d€ gylloid hum of Flt

REVIEW OF 

LITERATURE

CTIAPTER II 

REVIEW OF LITERATURE 

Th€ nost inDorta tuomnic rlrkr relaEd io $' Grylloid'a of the IndGPa* 

sub@nrirent and adj@.t @unrls e Gvh*d b.l*: 

Walker (1E69) @hlocued and d$ribcd the sP..imcB of Dermapt€ia-$ltatoro 

in llE collerio. of B.itislt Musum, along*ith kevs ro vriiou |ar' Out of 3o3 3p@i€s 

unde. 0. faoily Gryllidte, nt rcPoned lE sp€i.s hom the Indian sub@ndMt Knbv 

( 1906) published a synonynic etalogue of OrthoPl.n, mcnlioning 154 Semh 

undcr thc 

Thc crickcl Lrommv of undividcd ltdia atd ldj@nt @untris a mtumv 

arcnpr.d by chop.rd tn l9l, he srudicd rhc orlhop0cra of M4opotlFh a'd P'sia 

d*db.d $he OnnoDlln alongwith orh.t iB.cB ftom Ass aid Buru in 1924' 

idc ifi.d 6l rpcciB includisg 13 ncw tp@iLt of Grvuidt from C'vlon P@ it dE 

Brnhh Mr*!m in 195, Evisd the Indian Gryllid& in l92E 'nd published lbe 

Tri&crylid4 lrd Gtylli&. of C.ylon ii l 96 SatnE$gtn ( 1954) p@idcd t chat lisi 

of thc Trid&qtid& lnd Cryllidr. of Ccyloi ftntioninS 9t spei6 in 46 lptm 'nd 12 

subf.mili4 und.r thc l{Er. 

Abdukn (1955) colleEd 13 s!.chs of qict E f@n Lvtllpur (Faiel'!'d) 

Fernando (1957 a. b, c) d*ribil 5 iew lpcci6 of Gryllide fton @vlon Ghouri a'd 

McFdl!tu (1957a, b) studicd tnc reproductive islation and sp{i{ion in 'c"ro 

dont$i.B. rhr! d$ ery.imcnLd on thc developrcnt ot stttzt in erytud6

-5, 

J!it?d6 (r95?q l958a.b) in tlE l.bontory. r:dr a|d Asght (1951") tcryld,l Aaua 

paristdb ^tt Gt U.psis tdU4ddr6 at rew sP6i.s f@n LyallP!. (Flielabdd), whil. 

Ch@ri and Ahn d (1959) ide illfd G.tnbpsis pa\isroao fd $c l6t tine frod Mdn 

Chop.td (1959, 1960, 1962) studied thc srylliG of Id atd Afghanithn 

llongwirh thG of Angol.. In 1961, hc divided lhe 8.nus Ot {tr on Oe bsh of 

cogulnory organs and shiftld l0 Indian sp€iB ro his rcw ge.us TeLotry!tut Mali 

eid Or€m. ( l%3). McFdl@ ( 1964) and Mdhad ad McFdlaE { l%E) sMi.d vsio!3 

f&torr.ff.ctin8 t. srowth ad wins developmc in crickeli. Rddcll (1964) revis.d 

th. tubfuily Cryllire o. d. bsis of mle g.niblia and arldcd orc t ib., oE sutsdbe 

and 6 n w screo. Wdkq (1967) rcvised $e subramily O@nthi@ of Arerica. 

Bhownit ( 196?, 196E, 1970, I Cll , lvls, I qr6) studied th. c.icLr fauE of Indi., Nofi 

B.ngal,.t.,.nd dsibed s.vcral new rp.cics frcm this Egion. Cha&d (196?) 

crialosu€d 7E6 spcci* in 100 gcfta and 4 Fib* under th€ subfaiily Cryuina. of rhc 

rcrld, ncntioninS ll 3pccies fton Pati!€r. Ii 1966, h. de etalogud 1416 spels 

and 20J gerca wd.. 9 subfuilia of Cryllid& alongwirh O.enhide and 

Cryllodpida" of ti. wld, Siving only 3 +ei4 ftoD Pttbtrn. In1 9. lE tuil.r 

monoglrphed 355 spe.ics and E8 gen R in 12 fmiliB of Crylloidca from Indi2 and 

adjc @unt a. re?onins only 14 3pei6 ircludins 5 ew oB ftom P*islan. 

Tandon. l Shbhodia (19?2, 1974) dlle.ted 22 speie (imluding 3 new oB) arom 

Northst Fronri.. As.rcy and d*ribed 2 rcw iF.id of v.lanidow trcn 

D.hE Dun ad UdeQ6desn i. Indh, Vblc.y (19?3) ditgn6.d a n.v Cens, 

Dianmbius, ot th. etbfmiry N€nobiinae *i$ En@ptcft f@ips .t ttg tW. 

sp.cier. VMth a al. (n5) îA Venrh (1978) idcndfi€d 3 ncw ipei6, 

vel4rtf.tow jaiktiatLt, v. kh8i.Bis îd T.Itoerytiut Mqhalay@B ftoh 

M.gh.ltya, India and 3 mw f.mals ol V. j4iniM, v, klbsi.Bis ad 

St.phobLw hohb.niells &om Nonhsr India r.Ep@rively- coreknov (t9?8) 

worke4 on Tanatugryllls 'fa,n, ad Mdi.oeryU6 Ctop, of Fiddtc Ash.nd mrd. 

sonc i.aB6ornadoN. Kcvan ( 1980) sMicd thc orttopicreid ingrs of rhe B.rmuda lrd

-6 

clî6?dtjt:I Gry od.s swlkoLt h,r'jgtiotn, ovct c. ri8iltr6. TryMttd (l9el) 

clarifrcd O. @nfuion b.tf,@n two si'nilat Oricnirl sp€.iA, viz , Telzoe.rta dtatB 

Sunr.ister) ard T. Bnc.a (walrtt). H. prc?os.d the @. T eipiaut 

(Scdillc) rd rlE forcr and r drrarlt (Buroci.t r) for $.latEr' BlEsiq (l9El. 

f9E2) d.s.ribcd 2 rew sp@i.s, orc in a*n U.oN.6 ^d ttabt M itotr 

Nonh-w6t India. Corckhov (l9El ) rcvi.w.d th. 3ubfanilv N.nobii@ ol U S.S.R .nd 

t.atuf.rcd the sp4i6 h.M.tubi4 .tikii ^d P. taPtubMit b rk g.nls 

DioN@bi6 vi.t t\ ud alo.r€cEd a ncw genus &.@t.tu t6, In 19E3.h. 

d.*rib€d r group of 5 gercra rclattd ro Bnehliqp.t irclldiig 2 ne* 8erch !i2.. 

T.tbi6kiclll4 trd PlMr.UB wl:di, Acha. Ponattosaand 

rypd of ri. genera rBP@tively, 

etnrbSryu6 nn@r at 

TowMnd trs83, r€rB.d tn eeaos G+tbtabQ sith a lod or 12 sp;ics 

ircluding lhe 6 n* 06. h tni3 whion, hc.lto ovcrcd th. sP4is o' 4,!ro .nd 

c. on atr! fton rhe oricnal Esion. Ori! lnd Al.x.'d.r (1981) tuIbghPhcd 4q2 

spccis unds 85 seGn ar{ I I subf.miliB from A$ealia. Ore and Cad. (1983.. b: 

l9E4..b,c) and Oi& (l9e7a) err.Bivcly sodi.d thc Afiican hu6,nd decdb.d l?0 

sD.ci6 in U 8e|w, @utlom a dl. 09E?) dislsld ihe phcrctic ElatioB of 5 

s.nc6 of cryllid& fton Faisl.bed. Pqjni and Kalr. (198E) desdib.d 3 w ipe.ics of 

Tut6osryl!6, wi$ a rey to th. Indhn tp.ci€s. S..!d (1989) identifi.d 40 sp@i.!. 

in ludins 3 rew on s, in 17 gcrcra lndd ? Iimilis fion the Punjab (Pakttan). 

Alcxande. ( l99l ) evis€d the genus Crr{rr .nd rcpotutd 59 sPei€s rrcn difleEfl par6 

of rhc wold i@ludi.g Asia. Toms (1993) proltd in th. Iabodlory oEt Crylirdrt 

suqllco6 ^nd A ,iSt/l!r!J mre.4!!lly th. n.mpl. ous aid nicrotl.rcs rorns 

of on Nnd th. sc ap€.ics. Hc, th.rcfot!, syDonymi!.d th. laner enh rn. fomf-

MATERIATS 

AND 

METHODS

CHAPTER UI 

MATERIALS AND METHODS 

P.li!tstr li6 in th. nonh w6t of South Ali! dd .nends m t4orth fion 2:f ' 

l?' N lari$de and 6l' - 75' E longnudc tt hd ! tolal arq of aboul 796 096 squst' 

kilometeB. ofwhich about 59% formt high nounlains and hble land! in the mnh and 

w.!t. whil€ lhe rcoainins 4l 96 @Nist! ot sandv daens buren Placaus fenil' pldB 

in $c middl. alongwidr ccitl b.ach6 and n.rshv swanps in $c aouth TlE @ltiE|!d 

and fosl &6 @er 36S and oui4% of $. !o|al tr* resFclivclv Fiv'nainrivcu' 

i.c.. Indus, Jbeltm, Chcnlb, Ravi a.d Sudcj, flow thrcugh n and atc rhc Min slrcr 

of inigation in th. cuhivat d Plais 

of fi€ Punjab aod sindh Nat!61 veg€radoo ircludB 

lough dry gf.s, s$ .tl buh6 and lN trs i! 16 dry dca, eldl' lhe mnhcin 

mounuiE @ coletld with woodcd fordr. 

As Pak6bn is locaEd on de nonn ofd. Tropic of CaGr' n hs a o0Limnttl 

rrpe o! clrlw chMcl.ri4d bv eroem. vdilLioB ol EmPeralure ln @v paB lh' 

slmn r tnpqarure may rcach !p o 5e c and diuhal qriation nay b' a nucn d 

I l" C !o l? C. Wintds dc @tl vifi mininun D.'n temFdurc of abou|f C in 

Janoarv. Nonhern mountlis e nuch @ldcr' whe@ d. t'nPc6or4 on Balwhisl4 

plateau are sonewtal hiShcr. 

P*nlan slffe* from a geftral deficiency of tainfall. Although th€ couotrv is in 

rh. noMn e8on. n B c\tEmelv dd. c\c.p( for the 3ou!h'rn 3lopa of Hrmlavs and 

7

-8- 

rh.3ub.mounhircu Ea.t, vhich h.ve a Binfall frod 76 b 127 cn. Balrchbt n is th. 

dri.sr p.rt of thc @unry pith e av€taBc .ain of 2l cm. w6Er. cyclonic dirorba@3, 

origin.ring in 6c nediiemnm, ale qus. an avcrage rainiall of 27 io ?6 cn in w.$.rn 

nounrai Nnd dr.n inmediaE foElyins 116, ehcrd n is Edw€d io 4 cn if, he 

plai6. A ldgc psn ofde preipiadon in rhe nodsn sou liN is in lhe fo.m ofsnos. 

Pa&islll ir adninni.ltirely divid.d inio ldr p@ifrs or PuriNb. Sindn, 

Bahchbi.n a'd Nonh Wd Frcnti... 

Aduh crickeB wcrc @lleccd duing dc y.!B l9l 10 1995 frem v{iou3 

lcaliriB (Map l) of thc fouowi.S iour climatic rcgioN ot Palisbn (Ahmad, l95Dl 

2. 

Makan Cos1, tab€la, Kulcli, Hydchlad dd Badin (nurpartcr). 

Sub-toDicrl ContiEdal tir Lln.ls: 

Khurd{, sibi. suklur, DGn lsmil Khan, D.a Chui Kh.n, qjrJpur, 

L.yyah (Thal), Midwali, Kh$hab, Muaffege[, Mullan, B.l.wdPor 

(Choli$an), Kharcwal, V.hdi, B.iaw.lragd, Palpatk!. Toba T.k 

Singh, JhdB, Fabahbad, Sargodha, Lrnore. SheikhuPuh, Kdur. 

Cujtuwal4 Sialkor Chakwal, Pawalpi0di, klanabad &d Pcshlwd. 

3. S[b-trcpkd Conlildld Uirb r&&l 

Mu@, Abbooab.d, MaMhn, Zidi S*at. Kalan, Koh.t, Ztot, 

Quet|!, Krrar atd P.r.,gur. 

4. Chogl -Kbdn Regio!: 

Th. specinens w* @llectcd ltom holes, godowN, laf litEr .M othcr 

d@ayirg ve!.t ble nar0er, lotting logs, bushc6, hedgs, crop fi€lds. e5p4ially at thc 

rimc of ir.isaion, srel ligh6, Iight uaps in fi€ ficlds, under sloN od ll.rt b.iks of

N 

tu** 

oft 

s$P 

.i,î 

{i 

)' 

5" 

ii. 

.:q 

. ia 

|^J 

6II.G 

p" 

dY 

M.p 1. Mlp ot PrLlru .boxbs ou..doo rFrftr.. or 

Grylhi&. 

a'9J.d 

'sr.dqn,'M 

29 Gtinnr. 4r. xd 

d. arA 

r2. kshd ( b 

.r h'jl|!

-tG 

watcr $urc4. Thcy 9w kilted in a cranid. bdrlc, pinn d atd th.n bodv p-t3 sd on 

a$ropriaE ettinS-ba.ds. TheHfter. $ct *rc prcp€rlv lelEu.d dd slold in 

collectiotr box6 baving Mphrhalem balk lrd co.9.x Powder fo. dFir sddy frcm 

diffete|i insect pcsls like dernestid bcetl.s, an(!, erc- These collccdons *ei€ als 

supplement d wid sPeimeN ol lbe pr.vious workcA s {ell s th. slod.nt!'coll.ction 

in rhc D€partment of Agriculturai Enbmology, Uriwsitv of Agricultuc' F.ir.l.bad 

Th. malc gennalia wrc diss€d by Elaring lnc $aituB cith.r wirh n'aD 

liom a *aicr bao or by pl&ing rlEn in a d6i@tor for lGl2 hou$ A loisindiEl 

mid-venrnl irEision was nade and tie.ntire conrcn$ of the tat iour or fiv. abdoniMl 

segmenb wcr€ removed, Tle spaimeN tcrc lightlv stufied wilh @tton and $e 

abdodimlqalls weE moulded &ound it, with O. hclp orBlue, to d rcar thtir oriSiml 

shaF 6 posribl€. Afiq cl@ing lhe.€nov.d viscn iD old l0% polaiun hvdrdidc 

for lGt2 hous. thc scniulia vgc t fs.t d out, $orcu8l v sh.d in diltilld wt'r 

for $vcral dM att pr@ed in glterim in hicroviats having dB emc nunber a $c 

sFcimcn fron which they *ee rcnov.d 

The idcntifi@tioN sere mde uing a evolvirg skge and a Wild M3B biNular 

micrcscope (10X x 6.4x, lox x l6x, IOX x 40X) uP to lhe sp@itic level' Thd 

nedurem.nh of v&ious body parls w.rc lakcn with a vernier nidG@Pc having a l6t 

coun! of O.0l dm. Thcn, th.n an8c, naB, slanddd deviatio6 and ntio3 wrc worLdt 

our. Thc fie. hand st tch6 of vdios talommkxllv importa p!ri! .ionswith thc 

geniblia of alnost all rhc sp6ies. *hcth.r Prcvioulv d6.ribcd or rcw, w.G d6sn' 

The diltibution of all dE de&mimd spei6 h6 bei mappcd out Th' 

charadcB of thc lamily grdp haec nol b..n 8iv.n in this Muscral duc io tncn a, 

availabiliry in litclalure. Ho*ver, kets for all the higher and lo*er tixa hlvc bcen 

consrucr.d !o m*e the luture identific.liod4siei New gemra dd spccies havc ben 

de$.ibed in dctsil, while onl! ll'u diif.cntial aid rew clEadeB if a,v ftom de 

published d6criptios hsc b€.n Sivci in c$c of tlrc.dv dsiH gcrc6 and specs'

ft. ritind., longlndr, dtin dc, rlld i|cr 6nprn!|t! |'d .|| d rwr{c 

pEcbintior fo. dl $. colLdim d!! tuv. rt o b..|| edd.d. 

Th! cl.!.ifl.|lh| rld tm$nobS, Gir. t) of |tE np.rtuily cryloid.. hv. 

b..6F.dyftllo*cdr|rlvabrR."dc[0961),Cttoo.rdC59)!dOn .dAtluldcr 

(19$) itr dir onrl|crb( All dE tyF T.cio.!! hN! b€.n d.po.i!d h |h. td 

Muxu. D?.rtEotof Agrkolod B'nodloSr, U.itt riryof AgdcdEq hi|!!.d, 

Att wi|.br urd h 6. Eri &! n ftlldt !: 

B.L. 

H.L. 

H.W. 

F.R-W. 

P.L 

P.P.W. 

E.L 

w.L. 

M.L. 

H.F.L. 

H.F.W. 

H.T.L. 

o.L. 

s.D. 

Eody lcrtllt 

Had lcolo 

tlad f,ldll 

Ada|||ll widb (ld r.$rd) 

A@ir Fmd|D r n 

Pat.th. prorum widrlt 

Elytrr lcn!{r 

Hird sia hah 

Mirq lcnl 

Hlrd ftou h!gr! 

Hlrd ribir lcrgih 

OviF.ib. l.jdh

t*" 

I 

tu.r. 

-12- 

Pri'dp.l norphototlcd f..trG of I ctklt€t'

RESULTS 

AND 

DrscussloNs

CHAPTER IV 

RESULTS AND DISCUSSION 

Thc prcs.nt inv6rigltion rcgrding thc 6icLr fau of P!ti!'rn ha 'i:tlv vrcldcd 

6E spccica and subsP.cits undd rhc slF anilv Crvlloid... Thcs' lp'cica :re $nngcd 

in 2E scncn udd E fa0ilks. 

K.Y to rh. fullL. ol Gltllold.. 

l. AnErid lcSt .nl$g.d, n$h !@r.r trn niddl. !P@idi"d fd diS8itg 

(Fi8. 2A)i c{iPdior sb$nt. . . . 

Ancnor let! nol .nlse!d, Eidd tio{E thrn middk M 

diggil!; ovip6iio. P@n|. 

2. s@nd |.EI egmcnr hc.nh.p.d doB+Enrillv d'?B!d (Fi8 2D. with 

Premimnr adh6is Pad!. 

Se.ond mrsel !.srcnr 3|!dl, IaLEllv @nprci!.d (Fig. 2G). w now Prohinc 

3. P6Erio. ribi.. dtrEd *ith M. $at 3 lPitr.s on @h tna8in' scrrul'r'd d bcs' 

and beift.n th! 3piEe (Fis 2B); aPicd sPuls 6 (3 dcmd. 3 i d l): 

ovipodibr cyl indrhal, E{rly 3cii8hr. ... ... . h'oP'sli'|.

-14- 

o 

/7> 

t/" 

GrJlolde. A,.ndr L., c,rnd,Ir. toLttu q. ar 

(Grr&.ddedr 3, pddld dbL, L,qd!. 'zM 

(E6piatrb); c, rL. @, an4t tu ,rntlt q. 6f. 

(ftgddrlfdr} D, th mq O..N6B .hr'gbt sp. Mq 

(odnbtndr r, tf,. na Ob.dtr rd'iw dt *. @r. 

(Molo,||rn.dr r,9o.dort rE L rwe(F !.a&l; 

G, t|. r.a caitr ,r@d!rd (&tB&.' s, !..r!rd b6.l 

cj.t, o. .dnotlut $, m'. loa.dhr&.I li tL llft, 

n.Ioltuohn e&tu q.N. (Pll|!roIddx).

P6l.rior ribia€ wirhow spin€s (Fisr. 28.66D). 

Pdterior ribie provided wnh spin6 (Fi8s. 2D. 3CD), 

5. Eody more or l$s mvercd wilh scalcsi clyp.us .xtnding oo lo the doBum of 

6, 

1, 

t. 

m!.gi., non-smlar.d at ba. and 

(3 cxte.Dl, 2 iote.ml): ovipoirot 

Trlsotridlldrc 

h€ad (Fis.65A): f.n ld sinsl6, nd6 Gudlt enh sltrcvd@ clyra. 

Body glabrou or fe.bly pub€sn1, but eithout sls; clr?eus not.xEnding od 

!o the do*ud of h.ad (Fig, 55B)t borh exe3 widi tully d€Elop.d clyt6, , , . 

without ciiulation at bdc and 

spia (Fi8i. 3CD). Cr!lll&. 

P6krior ibDe seiiulaEd at bM and beN*n the spics, 

Slender msrB $r$ head lenSrhened. progurhous:elyta drn. 

das bifu@t (Fie.2H). 

Robusr insc$ with hcad 

brsal das no1 bif!@tc (Fic. 2D. 

Family Gryllidae 

progmrhoBi elyrra rhick, 

k is Gpr@ntld by 52 speics undq lE ge@rl and 2 3ubfamili6. 

Key to lh. subfrnili.s of Crtllid.. 

H€d without doaal brisdcs (Fis- 3A)t p6tcrior tib@c 4me

Ilg. 3. 

D 

.aP 

G.}u.la. had &d !6tu. t.id. daJti.,r.d 

{Grilnlr A,tt.'a ^, P&,u.tot/I. d..o&' lN@i||di; 

c,tr'n ,J,t4 calbt ,te'/,'s lcr'||ld); D. rh' l.& 

2 d.,rr. lNoblrudr E, ndbv pip!' C D,e.,.d' 

(G.rutu Or t,lh.tn, /'. @rol@ (N@b|rod

2, 

3. 

11- 

H€ad wirh ddel b.i!d6 (Fie. 3B): pGarior t'bi& 

spiB (Fie. 3D); mdillary 0alpi *ith l.sr egNnt 

prc.€dins o* (Fi8. lF). 

Subfamily Gryllinae 

dmed *ith long, pub*nt 

at l4r twice 4 long a thc 

It is iepreented by 49spdi.s ond.r 16 8.ncE in lhis cgDn, 

K.t to lh€ t€n.m ol Gryllin@ 

Oelli miged in a nqdy stRisht line (Fi8. 4A)r posl€rior fenora Bually d 

long d dbhe .nd |!rsi toge$cl. . . . . . . . . . . . . . . . . . . . . . . 2 

ocelli dBngcd in an obrls fianglc (Fig, 7A); Pdteiot femo6 rct Ionger thai 

tibia€ and bdirarsitogeth.r, ... . .l 

Body srolt and ldg. (30 nm)r p.orcruB much.nldged in fronti ovip6itor *.y 

shrt, 0.3 tin6 s tong 4 pcLrior Lmon. . . . luradn&s CorDLh.v 

Body than lrd snall (15 mn): Pomtum m! enldg.d in ftont dip6ito. 

modeat ly long. 0.? rimB a long Nt pdt rior f.mn. 

Cenialia wit virga vely 

elytra, iario of iB length 

sho.t, api€lly brdd (Fig, 9B): iemal6 with shon elyta. 

rhar oi ovipofilor lc$ dan 0.3O (eacrtt in Tuwoerrlllr 

long, apicrlly Miiow (Fis. l3c)i fenales with 

lo tbar of ovipNilor no.e dBn 0.55 {€xePt in 

Femde elyFa modqatly lug. (l.ll . 3.26 tim* a long 

overtappi.g or Ery narrcwly scparatDd on m.did liE 

€piphzllB u'rh 3ry|| on pscirq llicr.l lobe (Fig.9A). 

loDg 

6 

(Fige. ?C, 6). Male

5. 

L 

9, 

,18. 

Femak clr'tra very small (0.38 ' 0.46 timd a lons a 

s.petcd on nEdim lic (i.e-, 1.6 - 2.0 tims of d.ir lolal lcnsth) (Fis. llB). 

Male epiphallus witout styli on posl€rior laEral lob€s 

cylindriel ald la.Cc (l? nm)i wipGilor tong 

Body d.Dr6sed lnd lmall ( 14 mn); ovip6ibr shon (0.?5 

6. Eiph.llus with polt rior MgiD ml d..ply enargiEta. trEdian lobc a long 

or longd than latelal lobas (Fis. l2A). 

Eprphallus wiLh pdt4or @gin dcPly enwgin .. mcdtr lob.. if Prsnt. 

much shocr lhfl l&nl lot6 (Fi!s, 25A. 408) t! 

7. E lopdameres with bolh ante or pojecliois ne{rly eqlal, lpicls of th. fomct 

fi.g.Flilc (Fig. l2B)i body and tc8! altrGt glabrcc . . ot l,6 uNr*6 

_. Ectopahmcrs wi$ incnal anleid prejedoN longo $d th..xcmlom3 

(Fig. l3C), rpicd of rhe former vdiabl., but nor lingcr_like; bodv and legs norc 

or less Dlbdsnt, , , . , , . . , , . - . . . , . . . . . ... .. E 

Endopaa'iers C*hapcd tFig. 20D): mMl lob6 vhh their ati45 b.$cn 

apGs of rhe edoparincr.s (Fie. llc), 

€ndopuameres J.shapcd {Fig. 22E); mcaal lobes *ith rhe( apies belwen 

b8s or o.4!opdm6 (Fis. 24D). 

EpiplElhs endiog ina single elong.te media. prGss 

v.ry long and wto* (Fis. l3C): nircr 0.93 - l.3E 

ll 

(F'g. l3B)ralopams* 

tins6longswide... 

rilobar or roughly quaddte (Figr, l9E, 208): 

and br@d (Fig. 20C); nhror 0.?7 _ 0.89 tins a 

l0 

10. Epiphallus agically trilotst , with.nt rid odEin daplyemaGiuc (Fie. l8^)i 

9

l9- 

eodopalaneB wnh a rtuv.M paanclal must. apodene (Fi8. l8C). . . _ . 

.,,,.,,,,,...... ptartq4 u, C\opsrd 

-. Epiphallus roughty qudlaD, wideniog posrerio.ly, wm anEflor mrgrn nol 

de.ply cndgiMte (lie. 208)r cndoparanc€s withour a r.amvee pramcr.l 

muscre apodeme (Fig.2oD). pndloroboru, E n.nor, 

lr. H€ad qith re wid.. pate, raNvee banls on do6um; prcno$m febty 

widcninS in front, iB tal.Bl lob.s y.low, wirh a side blwn bad ato.t suF.iq 

@8'ni c.topMn 16 vitn ' €rDtanicdor prcj@tiotu origimdng cl@ ro thci, 

bd.s(Fis.22D). ........_A.t.rdFrbrtclus 

-. Hcad wirhou any orMn€dationr promrum dhdnclly Mro*ing in f.mnr, irs 

lalc.rl lob.s {holly blackt etoparah.tB widj internal mErior prcjelionr 

ori8inarins cle lo rhen api6 (Fis. 24D). 

12. Ant rior tibiac previded wir[. sioglc out r 

-. Anr.rior rjbie p@vi&d with horh i.Er $d 

13. Fiontal rosrtum ndow, $ *idc as or l4s rhan lst mtenDl segn€ntr clJ,p€F 

flonr.l suore very t@bly angutareicpiphaltu! wi$ b.dd p6lefior emarginrion, 

slighrly norhed ar middle tF18. 2tA). 

Frcnlal msrun widc, ducn wjder dEn lst anEnnal segne : cryp4rrcnhl 

suru.. stongry angula,re, turied oDlo rtE dotsun of had: epiphaltu wnh dcp 

a vcry drow pct .ior c@8iurion, rirhour a nedid ml.h {Fig. 2?A). 

Tsrummut! 'ftrblBktl 

14, Face lrongly ilauened, esp€.ialy in 

prcjering berween $e airnue (Fig. 

ow€r pa of dre fronEl r6trum. 

tl 

t4 

328): forehe.d in n.t. 

wnh anterior occltus on 

IdrrldM SiNun

15. 

-2ll' 

F..c |Er flltErEd, rFr. q lcss conwxi forcnad in nalc nol Prcjdi!|S b.t$.tn 

rh€ $r.nNt: fct|uL. tit|i anErior occllut on d|€ rpcr of fto .l l6tum. 

Ecbp.nm6 wit|l .Pic.l cxt rut drcn Produed 

t5 

inlo n ditllv sr.d' 

h@kdt rroes (Fi8.40C)i had totrctitrcs dor$Eninllt l.n8th.n d. . . 

, , , V.btilfr.bnt R.tiHl 

vrirbtc, rcr a .bovci h..d Mnd. nor .ro|$Etrr.dly 

Modicogtla C@ 

GdG Tt r&r*Ltur Cor.tl$v 

t983. Tabirski.Uts Gqotllov, Ent Obdr., 52(2): 314_330 

'f'tPc. Aclao Poacnoe Li.nwacin 

'Itc spccimcN of r siiSlc @lletcd sPei.s cEtlv t llv eith dE publilh'd 

dasrhdon (Cmkhov, 1983) of $b scnus 

L Totti,.f,utt toti.t&&r (Llc!t'r'tdn) 

17 . A.h.t Dott,toto LiclrcNcin. C.t. Mu!. @1. Hrnbu4' 3: E5' 

l3].1. eu ts Acha &tarie Sloll, SPcfts Grvll AclEr" p 4. 

tSt3. GrjAB (Adtdo) luDsiaotut Stoll, Sp..E6. Crvll Aclt t.' P 5 

rElg. Btuhy,tod !l,!&l&t S.dillc. lns. Onh p 326 

tE39. Gtr ts uhtliet 8uftin r. Hudb. E 2: 735 

|\fl1. Ittuthtt'rpts ci.tr(' S{6uc, Mcn. Sjoc C.Fv€.25: l2l 

$

-21 

r9(.4. Bt@Dirp* pot rorLt Rrndcll, Crntd. Ent.,96(12): l5E7 

loa3- Totbi6kidl(s po.rat rr&i C,oroldlov, Ent. Ob@.. 62(2): 314-130. 

All th. lp4ihens gre$ly @nforn 1o 6e published dd.rrttion (ChA&d, 1959: 

Taodon and Shilhodll, l9r2i Sacd, 1989) of tti! sp.cics .rc€pt rh. n@mm.t3 or 

M€doNnent! (nm): ld 5! 

M.L 

12.2.16.1 

I d 5 9, u!L$.llcd, t*.o fm rh. s{d.nti @ll.dion in thc D.p.rurc of 

Agriculrural EntonoloSy, Univesily of A8riclhurc. Faisl.bad, Pakisnrn. 

A snrgb role hs be.n raorded for he nrst lime fon Prtista!, Earlis, Sae€d 

(1989) identifEd only a singlc fcmlc fton thL cgion, 

a_12 

'; 

5,O.

-22- 

GdB Ptot rttt!, CoroHtov 

l9a3. Phowlla Gdothov, Enr. Obozr., 62(2): 3l+330 

'tW.. GtMtttULt mtor ChoPard 

Tle coll@nd sp.cincB of a singlc sp€i4 Bmbl€ the publbhed d*ription 

(cookhov, 1983) or this B.nB. 

l. Phor.Nllu, tkot (Chopsrd) 

(Fi8s. 4A-E) 

1817. BMbtrtpg (qMo?ryttu.) .ryktuc.phatu (rec Seryille) saasm, Men. 

S@. ceNvc, 25: 130. 

1959. GrwEryuB del Chopdd, Slult8. S.ilr,z Naturk , P l. 

!9a1. PhowlUs dr.ot Corokhov, Etrt. Otoz., 62(2tt 314'X3o. 

Thc aFlineN mainly @nforn io thc Publilhcd deiPtion (Chopdd, 1959; 

sa.€d, l9E9) or thb ipeiB Gcpi in tl|c fouowing chrtut B: 

Sorc speallB uc .nln ly bl&k (i'rludinS thc had) lile the veictv 

tulow.dtt tt ol .ryntu.ph.l4t Ancn@*ithswitndtwhnlthcclMt' Apidl 

fi.ld of Dlc clyns witb tF6 rci'|! (fit. 18), Sc Bu.lly l-2 bdE!.d. Ctlitrlir 

(Fiss. 4 CDE). 

MNm.nts{|m):5d 59 

B.L. 

2.14 

233 

2.36 

s.D, 

t.s-g.a ,; .;,

-?x- 

tslamabarl I 9, E-vlt-93; Abbo&bid I d l4lx-qi chanla Mdga (Kdur) I 

I, l5-v!93: Faiel.bad I 9,51x-8?; I 9, l4_vl!89: cnjd (F iola!'d) 2 d 3 9' 2& 

lx-88i I d 2 9,29jx-88i Llyyalt I d, t0tv-89i I ?.25'Vl-69i bnoe 3 9. 8'Vll 

89i Pakpa$n I 9,9-vti-88r I d 9 9, l2-X"EEt seeodha I 9,5_x_88i I 9, Gx_88i 

Hqd Sulcmaoki (Bahawalnagd) I d, l4_Vl'93; I I d l8 ? unlabell€d lale' fron the 

sr!dcn6' cou.ctio. in O. D.tantu of Agriculnirat Enionologv Univ€Bnv or 

Agricuhu.e. FaBalab.d, P*islan. 

This sp@ies bs bee coll€€d fron tiSht rcar cllrivated d'd of sug@n' aM 

vegeiables and f@m grsland3, herbs and thc fo..n planation 

lr ir rdd.d for th. fiN tin ftom Abborrbad Chans. Mdga" hlsntbad' 

Lallore and Llyyan. Eqdi.t, Chopdd (1969) lcpon d it fon Lvallplr (Faielabrd) $d 

Rawllpindi, *hile s@d (19E9) frcm Faisalabad. cojB PakPattan. RaMlpindi 'nd 

SBd (1989) pl@d sne speimens of fiis sp@i.s under OMo8ryl'6 

eDfuoc.PhalA on tlF bdis of anternae which aE *ithout a whitish ring Tlris 

ch.racre.. ho*ver, seeru to be ircoNtrnt d the preseot @llectioh conla'B |he 

sp€cim.8 whicn rc {id of wilhour a whitish ri.g in th'ir a 'n@ h n'v be rurthcr 

add.d rhar tn. odurcncft ofviious bodv P6.t5 in bolh ivpd of 3peim'6 Bradv

0.1b0 

l|.. 

I 

t+ 

A...rb i.!.. nr ar l ira 6. oa arE tr 

^,, 

c' *L. bl.r.4 D'1, *t...rb, r-l I 

I 

D

diffd frcm 0Dseof e. er:/thtoeephalnt, bnreacny tally qith those of G diiri It 

was, rh.refof!, thought appro?riac to pl4e bolh typca of sp.cin s uldq tt. l.r|.r 

G. ertth@ephtlla is coNidced to b. a sriou3 pdt ol semieting @uon 

(Yunus.,aL, 1980) in Pardstai. Act\ tty. it ts PhoMNllB rr'e., b@us the former 

sFci.s wa ncnhd found in llcld nor in thc nuscun spe.imns. P.ftap!, n Ei8;ht d 

Ge!6 lurorolrrlz3 TdblNkli 

1944. TrntogryIla T{biBkii, Salr. Ortnopr. IB. A4rb, S,S,R., pp, 19, ll5. 

l J. Pot4gtyllopsts Chopdd. Bull. R.c. (@m..ls@l llB/;,p. 169. 

T'tpet GryULt l4t.ruli. Fi.h.l 

Tt€ coll@Ed lFcineB of fiv. specics exrcdy rally with rh. poblhh.d 

d*ription (Ol!c, 198?a) of thb gen6 .xept dE clyha, which arc 1.0? - 3.64 tiru a 

Ket to lle spels of TMogrrU4t f abllsU,n 

l. Elrta and win$ @iplellly d.vcloc.d in bod s.x6, thc fo.nc. mr $an 3 

iims a long $ pm@om, $d,r.drd lrtlt.t Asgbd 

_. Elyll! @nplctcly devcl@ only i. nalcs. nuch cdlc.d i. ftftal*, l4t rhan 

1.5 tihd a lons a pronorun in lhe hicr: win$ 6uch rdk d in 

boln s.x.s.,.,.. -....,...........,,.....,.,.,,.,,., 2 

2. Occiplt with lo.g. prohiftnr ycuow lims; prcmrun }tllowkh, wnh a dek 

brwn h.dian bdl dd. few brcwn spolr E{ pdErior tugin. .. -,.. ! 

_. O@ipu whh shon, dull yellowish lindi pronorum brownhh-black. wirh llighny 

liglrt pyrifo.m impr6ioN. . . ..........,...,. ---. -. -.... 4 

3. Promtun mt @inSuariorlyi minq 1.53 

6 lonti .ly[a in f€nalc ndowly sepeared on 

- 1.56(Dd 1.55) iBa si

-26- 

Pronotud 3trorgly nairowine ant€riorlyr minot 1,67 - 2.0 

wide a long: eltu_a in fcmale dc.latping on ncdian 

(Fis.?C)....... 

4. Prcm$m bulging at niddle, not nalNinS anleriorly, L3? - 1.46times as wide 

s longtclyp@-frontal suture srongly angulaE. .. -.. -. ,ltlali sp. oov. 

_. Pm.orun mi bul8ing al middle. ndowing at riorly. l 52 - 1.6 dncs d *idc 

N longr clyp@frookl sutue sliShlly angllale. . , , . . , . . p@dtc4t sp. nov. 

l. Iumnomuus rsrirx.r,irj Lldf ., A5gb0 

(Fis. 5) 

1957. cryUopti! rslitr.4rd Llif a Asgnd, P@, Pakist. Sc. Cont, 81 59, 

t 3. Pansttllopsis rerl,n.drrr Chopad, Bull. Rcs. Counc. hr&l ll B/4, p. 169. 

1 7. Tuonogryllu rdlir.arid Choprd, Orth. c:r., pd 10, Gryll., p, 36- 

Th€ coll4tcd mtcrial la.Sely conforns to dE publishcd desfipdon (Ltif.od 

Asghar, 1957; Chopard. 1969; Sacd, l9E9) of thn sp@ier, but diff6 in th. following 

Prcno$n wirh poscrior m&gin sligitly sinucr elyfa in mald with ninor 

dividcd dighdy b.hind lhe middk (Fig. 5)r api@l l'leld having 6 - E veiN: latlral rr€ld 

pr4enting 6 - 7 veiB, Sc2 3 branchcd. 

Meaurtmnts {m): Sd 59 

3.G3,t 

Mrle 

s.D. s.D,

-27- 

hlamabad 7 d 20 9, 8 Vll-93t R.walpiodi I d, 8-Vll_EE: I d, !5_VIII-6EI I d 

I 9.25-vlll 88r I d, l-lx-6Ei I d,2Jx-88i I d, 3lx-68r I 9, tlx-88: I d.8'IX_ 

88r I d, gtx-8Ei I 9. lGlX-88: Fsisslabad I 9, l9.Vf93; Palealh I d. E-Vll-88; 

I I, qvll-88t Muafidg&h I e,20-VIl-88r Miarwlli I 9,4-X-88i kyyan I 9,29-V 

88. 

Th. ip@imN of &i! s!€.ies wcrc found in $c hilen l@v6 ad l@flin r unds 

deis plan|atioN of suflida (Ercalrptrc cMldukntis) a.d on gr.ss6.€,al light 

lr h6 bcn r@rdcd for thc fi^t dmc lrom hlatilbad. Elrlier, hifud Alcl'e 

( 1957) l@d.d it frco Fiisalabad lnd sa..d (1989) fron all O. abd. lqlili.s. 

Chopard (1967) dcntio*d his Mm 6 lhe origilul a!$or, eho d.sib.d thir 

speies in 1963. But a $c sne speci* had prcvio$ly bcen describ.d in Pathtan by 

klif and AsBhr (1957), thc priorily gH b $eo in being thc origiml au$oB of lhis 

Th. spdid.N olle.ted ar high altitude are da*d ii geenl colouraiotr and 

noE sPortcd al protoom than tho6! from Oc plains- 

1.23 

1.21 

0.21

-2V 

rb, t ,}tst ttrAat ,rs,4.,,t *b.3,

:29- 

2. Turo'towuut pa*idaad Chourt.t AbMd 

(Fig. 6) 

1959. GrtUopnt p64kkau Gh@ti., Ahmad, Pc. P.kisl Sc Corf LinoE, Il:48. 

1963, Para|ryuoptit r'o*liarA ChoPld, Eull R€s. CouN. htul ll B/4, p. 169. 

1.N1. T!runo|ryl\ls zalbkrst chopard, o h. c.t, pa6 10, Grvll., P 36 

h g@dy agces wnh rhe Publhhed des.ription (Ghouri and Ahn€d, 1959r 

Chopdd, l%9: S@d, 1989) of this spccies exc€plthc followi.g ch@teB: 

Elytra in malc! wid nitor 1,53 - 1,56 tims s lone d widc, h.rp! wilh 3 

oblique vcim (vsy nroly 2), lpical ficld wiu 5 '6 vcint' lll.al fEld with 6 _ ? veiis 

in mle and 5 - 6 in f!6418, S. bc.ring 2 _ 3 bhrchd. 

M6u.emnts (m)r 5d 29 

Md. 

r2.8 

306 

Faislabad 2 d 19, l2-vl-88r 

05 

0.l l 

,.;., 

"," 

I d | 9, l4.vU.EE; IslatEbad 2 d. E-VI|-91, 

.;

-3! 

rt,6 ?lJ[!''e,,|'/e|!, ha rt-ra Qt I,

-31- 

Thc apeir.B *ec collccEd in rhc lalliilcr ard oi gBs Ig . lighr $u@, 

It ha been @U@i.d for the flsr tinc frcm klanabad. Eirli€., Ghouri and 

Ahnad {1959) a.d Sac.d (19E9) l.dd.d n f.on Mdn (K{ehi) aln Failalabad 

sae€d (1989) ide.tifi.d on. fcnate 4s A. eirsukrw, but the prene of 

@Bpicuous longnudiml y€llow bands on @ipur, sliShtly sepdrd elyra &d ovipsnor 

wnh cbs.d apiol valvs rhows rhat it wa mjsideotificd and &tu.lly the sp€cinen wa 

The speimcns @ll@cd at high .ltiturh rc drtd in gcE6l otouRdon and 

noE sporEd at prcmtum and abdomcn rhd tl$6. collc.lrd in pl|iE, 

MALE: 

3. T4.6oeDAtt Lr64rtt 6P. mr. 

{Fi8!. ?A-C) 

siz. ncdiun (12-9 ' l5-5 nn). Body olouador brwn, tiig.d wiln r€ddi$. 

Rcddish-brcwr, dd6t .! wid. a .dErior sidrh of promtun; @ipur qirh 6 

very pronircnr, longirudiml ycuow lincs, inrcr two r@hing urnorly ro lle lalehl 

@Ui, tn. r.naining to the p60crior maBin of cyd, lhc m.did brwn band inte6a0ed 

by 6 very wrcw, loqiludin l ycllow lirct f@, mndibls and cn€€tl yellow, the lauer 

mlonr .xt€ndi.g through sidB of pdrclypcus up !o anterior Gll$; posicly?eus 

brcwnish at niddle below clypm-fro ial surure, rhe larter lrongly angulaE (fic 7A). 

Eye black. Palri yellowhh, 4tlt maxillary sho r than 3rd and 5th. A.teM. 

Pomtum 0.56 - 0.67 tin sa long a wid., distirctly ndrowing ir ionr, chi.ny 

in nals, its mr€rior Mgin slighdy @mav., p6r..io. alncr sFaight, both liftd with 

eddisb brtdB; dis* slighdy .oovex. y.llowbh, providcd *ith scar.Gd nre brbd6 and

. tw bo*n dot!, N bl.ckish-brcwtr ba d runrii8 in niddlc. broadcning toqards 

poslerior margii, wi$ a b.own sPoi on cilhq tidc, lar.al side brwn: lat€rd lob6 

Lrgs yellow, Pubeent; an€rior tibie pcrforard, sid r ldSc oval .xtlrnd ed 

a small. rounded int rnal rympanusr postcrior fchoa iliShdv adqncd enh browa sri.' 

on both fca. 2.83 - 3.39 time3.! loiS s wid.: PosL.ior tibi!. {ith 6 _ 7 in€rol ?_ 

8 exr..Ml lpiiB, th. fofi.t 3liShdy longcr lhan lanlr, borh @d.d and bownhh at 

aDic, ncdioinchals lonS{ $4 oilERr post rior b.si6si nodcraElv lo.8' fcblv 

@m083.d. .rn d with 5 - 6 inrtMl, ? _ E crEhll blowi sPiB 

Elyr. liSht brNn, AlMl trdpdcnr' 2 92 - 3.64 tiole a long 4 prcnoiun 

| .? - | .82 n6 s lonS s widc. tlidtly shofl.. d.i .bdom.n or @hins th. b$ts or 

€fci: nirq noch skler O.n long. t,5? - 2 O tire 5 widc s long. wirh ml! in and 

p6r.rio. .ngl* dMt ro!d.d, dividcd d@ io mkldlc bv u undul.cd €in (Fir. ?B): 

chords .adE.3[on8ly cu^td, lst uil.d wnh thc Firor bv orc vcinlct (r@lv 2); h'rP3 

with I .dly stdighr obl,q* vciN (vdy r@lv 4)i !pi6l fEld b@dlv reundeil .t aFx. 

brevided wi$ 5 veiB, cticulArion fom.d of n.rcw nr.8!lar @lls: ta.t 6l fcld wilh 

5 - 6 almct .Gguld v.iB, S. b.aing 2 . 3 borchB. WinSs nEh tllon r th.n clvlre' 

Elongal!. cylidriql, ycllowish wnh a b@d, n dian brryn band and a b'td 

ol $c eme @lour on ehhcr sidci epipral lon8. f!rceld. whh rceulrlv mdinS 

margiE, Cerci fltowirh, Pub*.nt. Gcnndia alnost similar to th@ ot T. paldttdB, 

with cpiphlllB d..pty notahcd itr tlic niddlc of th. p6t rior cMsination. laEhl lob€s 

providld whh long srtli, febly smll.n .p.xi doParaNG loiSd rhln stvl' 

deoricul.r.d Er b&, eilh. small r@th ttt ap.x on intehal Mgin' 

FEMALEI 

l! is samil- to n2lc, but dific6 in tic follding ch.fuLrt: 

Size l2.O - 16.5 nm. Bodv c.louo{on bl!.ki!h-brc*n. Prcnmd ld ttroiglv 

E owing in tront, qi$ rnor€ pronim tpo6 Ely!-a 3hort. I 2 ' I 45 timd long'r $an 

prooorum, .xEnrling a linle b.yotd 2nd abdoFinal scsn€ slighrlv o*rlaPPing on 

median lin (Fig, 7C), wilh intcrul oargin stoi8lv @nwx; dorsl fald ddk brown 

wnh nisd v€ins, 6 fr4 v.iB, Cu divid.di larral fi.ld v.llow[h, with 5 ' 6 pedlcl

tr.7. 

c 

-1! 

nwoEtls tal6.dt 

C ,h-4 pFDrm ud drbr 

{. oi. 4 ac d; E, dtf. ar 

B

-14. 

veiB, sc phin or turc.E WiDgs vcry !hon. Orhocilor long,0.E5 - Ll? tiG a lont 

s body, !l.nd.r, iG .pidl valvs smoorh, l!r@bc. er. ar !F . 

M.slrardrr (M): 5 d 59 

i,t.t 

2, 

E.6a 

s,D. 

0,,c 

0,31 

0_37 

"-,o.ro 

t" ol 

iolotr?€: I d, 6VIlt-93, Krlan. 

AllotJFr | 9, 5-VIlt-93, Krlm. 

P.rdylEs: KaLm 13 d 7 C, evttt-93i RlMlpidi | 9. tGVlI-93i p.tqod.n 

l 9, tcvtr-88. 

Tnc spc.imm lere collccti)d frcn tc tdfliler, udcr tlomr in ih. sit afil on 

ga$.s ne.! rh. lighr sol|6,

Dlstlnguishing chortcteF: 

-35- 

It his ! grat resenblan@ io t palir,ar6, frcn wltich ir can 6ily bc 

separarcd by i6 pronorum, which isdistinctly Mtwing in front, pdticuldly in ml.s, 

dirk hskinls, nirro nuch wid.r dan lonS and slichuy ovdlatping clyta in lhc 

tl5 nnor sho*s its afiinity with l. irti4 but diffeN fiom il in the long.d 

prcnidnt @ipit l liH, d.riorly Mding pre@tun. 3 hdp veiN. apk l fi.ld wi'n 

5 wins a.d diBhdy ovdlapping.ly[a in th. fcmal4, 

Th. name oi $is sp€ci4 hd b.en dcivld ftod Kalam, i(5 O?€ l@lily. 

The type nallrial hd ben deposited ir the l6at Mueud, Department oi 

Agriclhulal Enomology, Univcsity of Agriculture. Faisalabad, PakisBn. 

MALE: 

4. rtumuut bullaN sp. n@. 

(Fist- 8A_D. 9A_C) 

Si4 tudi{m {14-O nn). Sody.olounrion bl&kish-bown, wi|h lcgs vcuowish. 

Blackish-brcwn, shining, slighrly narower fian mdinum widot ot P.omtumi 

occipurprovided {ith 6longiludinaldullycllow lifts and avery ndrow obtue nedEn 

liDci facc, cheeks and nardibles yellow, lattcr @louradon of $e forder cxtcnding uP 

o arrcrior oc€llus; clyp@ffotrhl suture strongly angulate (Fig. EA). @hin8 up ro 

anr.onal skcB. Latlnl @lli ringed wih yello*. the ltler @lour exEndiog thtough 

a cnn.l sekcte i.to rh. y.,low @lour of th! f@. Ey6 bletish-b.osn. Palpi vcllo*, 

Anr.n@ yellowbh brown. 

Pronotlm stongty devclopcd, bulgin8 in middle. with anterior nrrgin f..blv 

comavc, posr.rior aln6t staighr or sligbtly sinuLi dhkconvd, bl&kish-brown. wnh 

a little lighter pyriforn inprasioB. provid.d with snall brislles and silky dark 

pubeseicei lateral lob€s yellow, with extrcmc irrterior ndgin alnGt sEaight, sliShdt

Itt. E. 

'37' 

aarding p6t riorly. lined {ith brown. 

trgr ycllowish. pubcs , widr b@nist hat, all fenoa t€eblv ring.d with 

brcwn at apist an€rior libi& eith a lugc, o6t cxt rEl and a sEdl. d'd€d incral 

post rior ftdo6 sttiaEd sirn btosn on both f&6, 3.13 tioB a long a 

rynpanuo: 

wide: p61erid tibiac 0-?6 tiG 6 long $ fcnora, emcd sid 6 idLrnal, 6 qErMl 

spind, ddkened ar bsB ard rPis, medioinl.rn l ,piql sPuB a long a or digndv 

longer than superios; p6t rior bdirarsi .lon8atc, a litlle mnprsed prolidcd wilh 5 

i €rnal, 8 external small sp'n$. 

Elym dlrk brcsn, transluccnt, 2.?3 tiFe! as long N prono$n r@chin8 to dle 

bes of cercir miror mdly oval, l .6l times 6 widc s long divided behind middl' by 

. cuN€d vein (Fig. SB): clod wid lst vein teBularlv rounded united wilh mirror by 

2 veinle6,2nd srongly curu.d,lrd ftarly stai8h! harps with 3 aln6t egulrobliquc 

veiNr apical fi€ld @owly rcund.d at.pcx, pmvid.d tirh 6 vei6 tnd rcticulttion 

rbrmcd of ndrow almd rcgulu cltsi lacnl icld vellow, msPaftnt rts sup'nor 

m&gin lim

-l& 

r& 9. tr tr. A-C, '+E i', dodt, 

t

{nh 3 bndodilbl bh.l bt!t6, oE 

ycllowish-brcwn. Ovipotilor vcry lon8, slendcr, cu ed upwd, iB apicrl vdvar sndl, 

l.m6laE, na'lwly -u€ .r rpcx (Fi8. 

M.e|!Mrs (m): rd 59 

Mrh 

3,30 

2.' 

0.62 

-39. 

in middlc, otlEr ts d tolh si.lct. sE6um 

ED), 

12,5J4.0 

,.r*., 

l.t3-t,z| 

solotyF: I d, 23-Vlll-9r, ca.rg. M.ng! (Ksu). 

AllotyF: I 9. 23-vllt94, Clunga Mansa (K8ot. 

P|rltJD* kldabrd 5 9, ll-lx-94: MaN.hh | 9. l5"lx-9,a; Abbotritdd 

| 9, l4lx-9,1: Ptshayd. I 9, !Vlll-g1. 

Thd sp!.i@B of lhi! spcciB w. @lld.d frem rhc Ldlirrlr und.r dc@ 

fosl phtrtitioB ard on gr$!4 n@ . liShr $urc.. 

"; 

r.t3 

s.D, 

oa

Dlsriryun i4.narr.r6: 

4G 

lr hs acl* rcsemblanc€ b L r4hror6, bltdifi€s in ptomtun *nich is 

smnSly dd.lopcd, wilh i6 3id6 bulging at diddL, ditt bl&lish-brcwn. oc.,put eith 

doll lin.s, elyn in f.Mle vcty sholt and abortive wings in bo$ sexB 

Thb sp4is hss ben emcd aicr rn Mne of r6t famos sai and pat Bulhn 

Th€ rypc natcrirl hs ben d.p6nad io d. INdr Mus.um. D..p.nncnt of 

Agricultuol EnonoloSy, Univcuny of ASriculntr., Faielab.d, Palista.. 

5. T@omuat PtutEa 

(Figs. l0AB) 

sp. Nr. 

MALE: 

Sia smdl (12.0 13.5 nm). Body @louradon btownth-blacl wnh bss 

Dark brcwnish-blek, shininS, slighdy wid.. thd lnt .id widlfi of prcrcruo: 

@iput wi$ 6 re.bly lisiblc ycllo*kh liEs (vdy rely w.llnev.loped), ffi @ching 

rhe a .rior ocellusi f!@, ch@ks dd nandibles yellow, $e lan r colo( ds extendiiS 

op to rMrior @llui clyp@Jrenbl slNE sliShdy .ngulale, tut u.hing up io pot rior 

margin of anl.n@l sock€ts (Fig, l0A). Larcral @lli yellow Palpi t€llow. Antn@ 

brcwn, wid lst *8hcnt ttuow, sliehdy !n8.d with brownish. 

PrcmtlD 0,63 - 0.66 timd longd than widc, slighdy wrying eLriorly, wilh 

irs a lrior mdgin a lirtb @nqre, posterio. stnight, both ptovided eith small briltl$i 

disk black, virfi slighdy lighEr pyriforn iFpaioB, d.Bly Puba!4 ; htehl lob6 

yellow. with extiene inferior m&gin liied wilh btoqnish, eending pct iiorly, 

Lrgs tlllowhh; tll fcmoo sli8lttly hord€d *ith b.own at api€: rnErior tibi& 

wnh a hrg., oval .xE ml tympatum. intrnal3idc wid only a deprsion or a rcry

l|i lf, 

{l- 

n"\A VA^ 

o\J 

I 

W tt*t +-.Abdta.bd.

n inuc. reund.d orcl p6Erior f.nda strilt d with brcwi on both 3ids; pctcrio. libi& 

0.80 - 0-82 times 3 long 4 femoR, qirl 6 - ? inElml, ? external spines, the former 

nighdy lon8.r and sloucr Os lhe lalI.r, bo$ ddken d at bs and apis, 

medioinrelEl apiel spus s long as or slig\ly lonler than suPerointeroalsi posrcrior 

bsir.rsi elongat . febly onPres4d, atnEd wnh 6 - 7 i ..ul, ? _ 9 dternd denrhl6, 

Elyrra dzrk brown, lranslu@.t, 3.0E - 3.33 tim4 s loig a Ponotlm. 1.85 _ 2.0 

rims s long s wide, @hing th. aPi of abdomni mirc. oval, 0.63 - 0 70 riB s 

lons d qide, divided behind middlebya curu€d vcin (Fig l0B)rchords sttonglv b.nr. 

lst unn€d wnh niror by Mo Eiil.t!: harp wid 3 ohlique vciB; apical fEId plendng 

6 uoitormlynispscd v.iN, tcriculatioo foroed of r.guld, alnml squaE cclhi laGBl 

ii€ld pale yellowbh, mtrsparenq provid.d with 6 rciN, Sc 3-bnfhcd (@ly 2). 

Brcwn abov., yeuo*ish beMtb, with 3 f4bl. ddk bwn b.ndt oi doaun, ore 

in middle, orher rM on sid4. Cerci long, brown, pub*nt, C.nitalia very sinild !o 

$@ oi 7. ,!t ri sp, nov, 

M*lrt|mts {ml: 3 d

4J- 

3.03-3 33 I12 

0.614 70 0.66 0.01 

o30{-&2 0.32 0.01 

Holotyp.: I d, l4JX-94, Abborbbad. 

P&t.tp6: Abborabad I d, l+1X.94; MaN€tn I d, l5lx-94. 

fi. speimctr oa fiir sp@Ls E@ colLqcd fiom ln. ldI-litter and glls 

growo in a vcgerable fEld. 

Dislingunnlng .hamt r3l 

ThL 3Fcics h4 a cl@ @mbtz@ 6 T. kalMit 3p. w., but diffcB i" 

ib pronolal mking, dull @ipit l liM, cltt*frcnsl suturc lc$ aigulate, elyta 

llighdy @rowing anlcriorly a.d apacal rield with 6 veiB, 

Ir i5 ale cl6e lo t JalD..r6, blr @n 6ily be epdar.d by irs promcal 

markinSs, dull ccipital lifts, cl)?6-iromal suture less angulale and very sho wings, 

Thi! sp@ics hs ben Mmed, ar n ha ben d*.ib.d for the fiBr tihc, 

Tn ttF mat rial ha ben &pcir.d in $e l.gr Mueuh, [t€p.nm o{ 

Aglicll$ial Entomology, Univ.Biry of Agriculrure. Fabalabad, Pahisla!, 

G.nc Czlrdrlrlr Sjo6t dl 

t909. CalloqryllB Sjosr€dr, KilimandjeeMeru Exp.,31 105. 

'r,tyc: Callosryllu AAo@tdjonc6 sjxt!,il 

FEMALE: 

A sinele @llatcd female @mpl.Ely as@ wi$ publishcd dac prioi (Choprd, 

l. CalbgryUus or aneut sC, aor, 

(Fiss. ll ABC) 

SD. large (17.5 nn). Body stout, cylindricali colounrion palc tBhcmus_

Rounded, sbiiing black abole, 6 wide d prcDom; occiput with 4 obsolerc 

lines; frorr.l 6r!m 3,5 tincs s wide s lst drcnMljointr che.ki y.lld: fe Palcr 

dyp.cfrcmal sutuE sraigbt (Fig. l lA), obslet jn the niddle. Mdill.ry palpi Pal. 

bow., 5lh joini lon8.r thd 3rd, *idening aDd obliqrcly tru@r ar aFr. AnEn.re 

Prorctum cylindriml, L4 timd broad.r thrn long. i.r anrerior mtgin €ry febly 

concave, pdterior suaighl disk browrish, provided with snall rcddish-brown brbtls, 

slighrly @nv€x, wilh sides dn6r slaight; latehl lob.s palc whitih, *nh 3eaigh 

L.gs of tc eoe @lor e thc bodyt anl€rior iibjac wnb a ld8c, oval .nernal 

and a v.ry ninuc, rounded inEr@l tyrl}dusi pdterior femora Fli'cr short, srout. 

1.3 rim6 a loig a ribiae and 2.6 dh.s !s long d wid., pale wnh rcddbh tiisc on 

.xcrn l fac.i post rior ribi& wid 4-5 inErnal, 5{.xt rEl spioca: pcr.rior bailaRi 

wirh 4'5 irtenul, 6 ? exrcmaldent.ler. 

Elyna very shon, reduced b lateal pads, sepdat€d by a widc Bap on th. n€dirn 

lin (F'g. llB): dorel field rdaigllar, pre!.itn8 3 oblique v€iN: lal.Blficld with 4 

regularly spaced, alnost srraigln veins, Sc unbBnched. 

Ycllowish, gadually tapsirg postriody, with a wide medid browr band and 

a few spors of d. sd colo* loq.d larenl sides. Ovilosiro. v.ry long, alm6r d lotrg 

r\ lnxly, nctoiot. i.5 tines $ knrg as F{crntr tnora. will snall. miiow, acurc 

M.armedr (m): I 9 

B.L. H.W. P,L, P.W. tr.L, 

F,WJP.W P,LJIW, H,T,L./II,F.L 

g,F,L, H,F.W. n.T.L. O,L, 

Holoty!.: | ?, I vll-93. PalBEtr ForB' ra'i''k (P6haws).

Il& ll. 

r)'-' o 

VO OV 

45- 

\OP 

.CaIoSrtUt t rat]ngu q.6. fG e; s, h-4 Fde 

^, 

d dJi.. r, do.d .4dr c, G ..d ojr-.. 

t

h hs b*n @ll@r.d frcn $e le incr urd.r a dcne rost Pl.nuion. 

Diningulehltrg chrmcreB: 

A very dislncl speies Nnidr resmbl.s C Pall76, but 4n cdilv be 

disinguish.d by its la.ge and slout body, lery wide fronlal roslrun shining blrck hed 

wnh j6l a s$phion of lines on @iput and very long oviPositor (Fig. I I C). 

Th. Mmc of lhir tpeies hs b@n doivcd frch its ovipodit r, which b v.ry long- 

Thc holoryp. fetrule hs ben dePositcd itr th. Ingt M@!m, D.9|rtncnt of 

Ag.icuhural Entonology, Univerny of Agricultur, Faisalabad, Palhhn 

G€na C/t or'i,, ChoF.n 

ft17, erylods Sa@urc. Mem. Se. phvs. Hist El G.*ve.25:206' 

tnE. ery opsit Choped, Re- lnd Mus.,30: l3- 

T|l{f: Cryltula l.rr@&i sasuE 

A singl. tcmalc @tldtd entirclv confdEs o thc pnblish€d d4ciplion (Chopdd 

l%9: Saeed, l9E9) of $is geous. 

l. GryIoPtk Pub.tcent Chnqt d 

gla. eryUoptis Prrdr..d Chopaid. R6 lnd. Mus 30: ll 

h Bretly ttsbld lhe publtlEd d*dPtion of chopdd (1969) 

aMomiMl i.rgic. which d.ntrRty monl.d * h brM 

're$ r[' 

Mesum.nt! {m)i I I 

B.L. H,rv, P.L, PW EL En.L. HIW HTL OL' 

63 r05 

It,w./r.w, P,LJP.W- E.LJP L ll T L /tr-F.L lr r'L /o L 

0.933 oJt 0.161 0 650

AbborBbad r ?, ll-vll-93. 

47- 

lr 6 . @ry @ speics, ehich hd ben @ll.ct d fron ln. Pl..t d.lrir u'nc' 

v€ry Oick plankion alodg road side. 

h ha b..n t$d.d for the fiBt tim from Pakill,r' 

,'@ding ro CboPdd 0%9), it diffcA fion o$er meibcB of 6it SeNs in 

having snewhat de9es

"48- 

tE55- G'|UB lrytbrit Stal. Oefv. rct. Aled. Forh., 12:351. 

1869. AryUot itttinupt$ Walk€r, Cal. Demap. Saltat. &n. Mus., l:35. 

1869. Grylllt ruqi@rs wNlL.., Idcm. Ibid., l:25. 

l8n, Liogrylts ,teda16 sawurc, M.n- Soc- c.rcv., 25: 139. 

The @llet d narerid entircly conforms to the publbh€d dsciption (AHu[an. 

1955; Chopdrd, 1959; sa.€d, 1989) of t!! speica, bul difrcB fton otle .rd cadc 

(1984.) in tbe elyra, which e nore thm 3 tin$ a lon8 a prom$n lnd $. pcterior 

fcnm which @ 0.6ru.?91imd d long a elyur dd Lll - l,3O tiB d long al 

pGlerior ribire. G.nitalia (Fi8r, I2ABC). 

M6|lMEts(m):5d 59 

5,5-?.3 

1.21.1 

Mate 

s.D. 

0.33 

13.42 

''',

Ilr. lr. 

c 

4> 

W fu. r{.r.E r.

-50- 

hluabad I d I 9,8-VII-93| 2 !. ll-lx-94i Kalal l d,5nX,93; Zianr 6 d 2 

9, IOX-83; Zhob I 9, l3-x-93; Pa.jgu 2 d I 9, l5-X-9li Abbonbad I 9, l+lx-94: 

Pdhawd I d, l-vlll-93r Bhulbd (Murc.) I 9, l3lx-94: Fab.hbad I 9, l4-vll-E8i 

4 9, IGVII-88| Lahore 2 9, t5-VI,89i I d I 9, 25-VI-89i Murree I 9, I2-VII93; 

P3lpaMn I 9, 15 lV 89r I d, l+lv-E9i I 9, lE-lV-E9; Raerlpidi2 I, I4-VIII-88; 

I 9, l8-vlll-EE; Head SuLnanki (Bah.*almgd) 2 d I 9, l4'vl-93i Suktu I 9, 2l- 

X-93r Kdachi I ?, 2tx-93r l5 d 30 9, unlabell.!, ta&.D f@b ltudents' @lh.rion in 

$c De?dr!rcnt of ASricrlltuEl Enroml%y, Utrivsity of AEiolr!ft, F!i!:lab.d, 

Il ir no( @nno! and widely disfiblt d 3pej6 found in grsy .r.as, rcil 

ciacts, crop ficlds, &ound towB and villaga ad moc oft.nly @ll6r.d or ligh! 

Thb lpeica exlibit e@ @l@r vei2tioa, Nch a dE .lyF. @y la.k the 

chdac€rbtic ycllow spot!, 3hoa d e.rnely reddbh-ycllry @lou d thc l.gs my be 

partly reddhh or bl&kish. Atrotn r cmrkable vari.ry ir enrircly reldbh-brown wilh 

h hd bcn @dcd a a erious p6t of @non. millels, oil{4& ard shi! ar th. 

ntrBhing fl@B in D. G. Khd divkion and its adjoining al@ in Pltilhl (Ano.., 

l9l-?3;YunB a di., l9&). 

CerE r.&afrr[6 Chop&d 

l%t. T.koertlls Chopnd, Eo., M!dr., 3?: 2??. 

T}4ct err u Postiw \'t^tkq 

Thc .olleted mtc.ial SMdy tallid qirh plblhhed d*qiplion (Prdcu, l96ai 

One dd Alerander, 1983i Orl. and Cadc, 1983a) of drb geru .xc.pt the occiput shic[ 

is wiln or without yeuow liB a.d th. md!rcncnls of vdiou body pa.B, 

Ii is r.prsented by 5 spais ircluding 1he 4 new ond fron this rcgion,

3, 

-51- 

Key to th€ sp€cl6 of Tckogrtllts chopt d 

sid snall (bclow 16 nn)i posr.riot f.moia relalivel, lons Nnd 3bde. (le'gth 

ro wid6 ralio grcater IlBn 3.25)i sPiql field of eltra with 3 v.ins. . . . 

si4 l*g. {ahov. 20 m); pcErior r.llM elaiivclv sllorl a'|d 3to$ (l48th 

width .dio les ttd 3. t6)i .piol ficld of .lytra with 4 vciN 

2. Head with 4 ptodinent ycllov lifts on @ip!t; clw€cfronral surure st'odglv 

angulater laicral flcld of elyra pal;, wi|h a darL ba.d along it! superin Fargini 

8enr.l'a s rn figs. IIBCD. . . 

Hcad wilh 4 vcry fainlly visibl. tlllo* 

atmost st.aighri laEnl ficld of clyra 

fi$, L4BCD .,... 

liG oi ciput: cltp4ftond sutuc 

wholly ddk brown; geniiliz d in 

Aody @lour reddish-btowni h.:d with Pronirenl vellow bandt along inrcrMl 

margi, of.yB ttw also slcnrb b.low and b€nnd dE lar.r: laLral lobca ot 

prcno$n for $e m6t palt ycllow. . . . .. . . ,olrdl sp.nov. 

Body @lout blackbh brcw.i head withour dv mekinS or with fe€bly visiblc 

yellow ban& only alons internal naBii ofey* aid on occiputi l.teEllob6 of 

proootun lnlncly bl&kish_brovn .- ,. .- ,. ' 4 

Head {ithour .ny Mkins ircludiig thc occiputi dvm 3 long 4 atdomn (2.9 

tin6 6 long as premtun)i apicd ncld of elvra very long (1.65 tim a long 

s mircr). ... . ptu..tut sp. nof' 

Hdd witl r fe.ble band along rhe inl.rftl nsgin ofoch.vc alongwilh 6 rhort 

li6 on @ipul: .lytra slighly 3hottd tan aldontei (2 72 times 6 long a 

Dromtlm)i agicrl fi.ld ot.lyrB 3hort (1.29 dn6 a long a nnor)' 

.,,...- . .bngtidur 

2

MALE: 

L teloCryIus &h,r,r sp, tror, 

(Figs. l3A'D) 

Si@ rathcr lmau (13 14 nm). CeneEl @louhrion brownish-ycllow or broqn. 

Shiningjer bl&k occhur Nith 4 p@mirc.. bul short yellow lin6j .ye. blact, 

a@ behind ach yellowish. lalter @lour ,ko exrcnding narcwly alon8 $c intenal 

marSin of et6 up !o late.al @lli: he bl&kish-brow., with da.dibl6 r.ddbh-brcwn; 

clypecftonhl 3u$rc angulat4 Palpi for rlE nol pan p.l. wiilbh, 5rh sghcnt of 

naillary palpielongale, witl' apical half brcwr, slighdywidenin8and obliquely truncak 

ar apex, Anrco@ Dowo, 

ftonorum brcwn, more or les sport€d wilh ycllo*ish, sid.s brlging our medially, 

ant€raor odgin @Mve, pcErior fably sinuale, lirEd wirh ycllowish; larEl lob6 .lark 

browr in superio. part, dre emaioder y€llov, exr€n. i.f.rior nsgin black, asendine 

tf,gs ydlowish. noded wirh browi; anrerior ribia. wirh rynpdum oi inner a.d 

outer t&es, $e lbrFer shall, roundcd, rhc lauq ldgc, elongater pGErior fedora 

r.larively long, slender, darkencd at apies, a.med wnh oblique brcpn stri@. inrmp€d 

bt 2 longitudiod y.llow iripes: postdior tibi@ slighdy lhoner thd fchon, prcvidcd 

*ith 6 i.ner, ? outcr spins, l]]c fisr our€r r.latively sborr, nedioinr.rnal api@l spurs 

distirctly longs thln sup.roi.tcrnals; pdcrior baiBsi armcd wirh ? - 8 i.Er, l0 - I I 

Elyra brownnh-yellow, with light humeral bands, dcnding ro th. apex of 

abdon€n: Dirror wellJevelop.d, obliqudy l.igde..d, almost s long s qid€, roundcd 

ante.io.ly towards chords, divided behind &e middle by a cwed veini diqgonal vein 

nraieht !P to the anerior &dc of mnrcr, curu.d th.Hfi.r ar n@ly l@(Fie. l3A): 

c[ordr long, regul&ly cuped, very close ro diagonal vein; oblique v.iB 3, wcll- 

develoFd .nd sinuaEt apical ficld rcllievelo!.d, pKnring 3 vciE and rcricularion 

forn€d oi €loF almsr clongar. .@l&: lii..al fi.ld p,lc. r6Nluenr, eirh a ddk band

f,l& lJ. 

-53. 

rdottf,lz. IkNilt ry. d. '\ *rn a; DD, rd|. dt 

B, C!-tr-, d..l .a..tr c, D, !t|!. c4la, iid rd

-54' 

aloig supcrior rurSin, plwid.d whh 6, ngularly.pr..d wiB, S. b.arins 3 EiN 

(rftly 2). Wi.g3 3liddy longc. tlqn clytr.. 

Mod.d with bro$,n rd y.llowirn oi t.rg!m, iB licmun y.ll*ish, C!rci b|oM 

.t be, ycuqi3n ldstds alq. Cctriuli! (Figr. I3BCD) lEly snnih ro I. 

lorS?cnd, its.piphallw clonSlr€, cnding ii a 3ingle V{h6peit proccls, Brlting ftom 

a &!p natlw eDdginnion of rh. anErid nrrgiq dopffi clongaD ind fl6hy. 

Meu|tlmts (m): 2 d 

Holoatpe I d, lqvtrr,94, thasar (Toba Tck Si.gh). 

P.nt'p.: I d, |9-VIII-94, Bh.8r (1006 TGk singh). 

BoO tlE rypc slcciruN h.w ben @tk Ert iion rhc tdf-lir.r Ed ronirg togg 

in a dc@ fo6r pl$iirion. 

s,D.

Diningulshl4 cbrr.dN: 

This sp4ics is h0cl clm to T ttryiPantt, frcn ehich it diffeB in th' 

@louhdon ot lcgs,4 p.otnimnlycllow lin$on@iput, mitrcr altnoct.s lonS 4 widc. 

I obliquc veiB, lpic:l tield iho( with clos rcti.lhtion and glniral @nfisuratioo 

(Figs. 13 BCD). 

Tte M. of thb sp@ict has b€en dcdvcd fo.m thc f.ct thar both the couetd 

sPeircG w4e fould ri..r rofiing lo$ 

The holotypc and paratlpc of ihi! speiB h!v? ben dcp6iEd itr dtc l'gt 

Mus.um, Dcpanm..t of Asrbultuhl Enlonotogv Univ.Birv of Agricolorc. Fais'labad' 

2. T.Losrrutt lrn tilt,ti.i (SN,elit) 

(Fis5. l4A-D) 

t571. OTUB to.Sip.nnis Su3sure, Md So.. GctEv' 25: 16l. 

l r. T.Lo|ryll!.t longip.n"i! Ctnp.'4 ElA'rtAt.'37.2n 

Th. @lle.Gd 3P@imens of rhit spei.s SEdv agi!' with thc Ptblishcd 

d*riprion or Chopad (1969). bu! direr in dE followinS chaFcEB: 

15.61 lobcs of prcmluo with . Fllowilh spol at dtcrior dglB rh nArrwlv 

cxr.rds b4t alonS infcrior maryiE. Lces d&k rcddbh'b.Nn Elvt'i in tulc witl doMl 

n.d bdn at bas., larchl fi.ld dik btwn ficld wilh 3 Eic rd a otld 

'picd 

i.rc8uld rcdcul.tioni dyn in fcmlc *ith dqsrl fictd p@ndng 3 fre vci tnd 4 

hramh.t of Cu. wirg3 qud.t or vc.y tio( Ociitdia (Figt I4BCD). 

Mesurnenls (m): 2 d 29 

B.L. 

€,L. 

MrL 

3.30 

9.20 

slr, 

s.D. 

0.10

I|g, ra, 

.56 

reg4bt lo.gbatnt ^' dr|' dr &D, a-r.n a; 

x' #r+ d..rl +r C, D, !r.r. .qL, uld -a

f; 

2.11 

9.8r 

t.1n 

0,93 

0.8, 

-57- 

o,@ 

0,03 

0,01 

,0.J,0., 

oltt.t5 

L12-1,t6 

ro,3t 

ht!d!.d2 d 2 9, ll-Ix-g,.. 

ftb!c: 

ThG Np€cims of thb '' cr6 wcrc collaLd nm gare |'Idd r light !otE' 

MALEI 

of rh. body. 

Bed! 

Thi. !p.cl6 h6 bdn rccoftled br th. fiBl tioe fton Prkitirn' 

3. T.bogtllltt Jontt{,'i 

(Fist l5A-D) 

sP. tuv' 

Siz. Itrg. (2.16 - 26.0 m). gody coldntion ndditb+rom. L!! Fht tut tst 

R.ddlth-bwn, shirirS, tliShdv flrwcr thd pret)olln, wi$ r co$Picuout 

isrow }!llN bcid dong ilt rnd mllh ot cyd, qlending th.oqgh brlnl 6dli rlong 

dc alda3 of roltrm aDd rld ioltjn8 th! tn!*F v€lld bodl tj.bw ndi& oc'nB 

rhis band rl$ f:iltly dierd! h.kw ad b.hiid €..h .v.: fre !.lb*!ni dvP.o_fionDl 

t.1l 

0.0,1 

o03

-58- 

surur. srarghr. Eya yellowish, spon!.d wnh brcwn. Mdiuuv palPiv.llow nn !.8rc 

widcning. obliquely rundle and f.tbly da*.iitg ai apex Amenn& brown qnh cdg6 

of their s@ke$ blekish-brc{n 

PronooF duch widcr ihln lon8, slighdv *id.ning dEriorlv. with forc mdgio 

@nqve, hind sinrac, bofi prcvid:d with dcNe rcws or small brisd$i disk rcddisb' 

brown, d.6ely pubcsen, ehh 3na!l hair: lallrEl lob6 v.llow. *iti suFrior Mgin 

Mrcwly ddk b.own, ql'lme inferid md8in lined wilh bl&k pmdued psEriorlv 

irs anterior angl. alnost roundcd. 

L.gs pal. ycllowbh, pubenti 4lt"rior libiae providcd with I tvnpuum o' boo 

h@s, iind one small ad rcudcd, oul4l onc v€rv largc and @ali pcr'tior t?moh 6ick 

aid 3tour, 3.02 - 3.16 tines as lo.g d wide, f@blv sriabd wi0 brown on 'xErml 

sid6i port rior tibi& yellow, Ptovided with 5 -6 imr' 6_?out. tron8 aPlqllv 

cuNed, bmwi sPiEs: .pi.al spu.s long. mcdioit€rtal 0.71 tim es long a Pderid 

b4ilzrsus, tlF latt€r ptovided wi$ 3 _ 5 irrc., 5 _ ? outcr denttlB 

Elyt.a rcrching to th. ip of abdoncn ddL rcddish{own tdslwnt; mir'q 

loiger $an side, Ll4lin6 $ lotrg d wid. feblv dal divided slighllv h'tow middle 

by z sinlat v.in (Fig. l5A); diaSonal v.in v.rv loog and sraighi; chords long' feblv 

cun d, lsr uniLd *ilh diagoml vein bv 2 _ 4 veinlc6: hdps witn 3 _ 4 min, oblique 

sinu& veiN a.d ofleo Nith I _ 2 f@bly vitiblc ssrllcr orcsi 'Piql fi'ld !ho't' 

pr€senri|lg 4 vei6 and EricularioD rormed of.erlv elongatc @lbi latcral field blown' 

lnh lO- 14 fdbly bd.egllulv{paed vcic ldt 4 ' 6 beins br@h6 of Sc Wing3 

very loflg, erending wll byond the elvtd 

Bl&kish-brNn above, ycuowish b.@ln. c€rci long vellowish'bto*i Ccni|alia 

(Figs.l5BCD) vi$ .Piphdlus .tonsaE, fomed ofa.inslc pcErior pi@ wirh 

' ddp 

a erior cnsginadon and sinuak larenl sidca. is lip gnduallv Mrow'd forminS I blunl 

medirn Poi.t. 

FEMALE: 

with 4ropmn rB €longat ad n*hv 

It rallis *ith th€ nal. cxccpt ih. followi.g chd&t 6:

Ft. ls. 

DC 

.59- 

t riofltllat Nantll ?. Er r\ dttt dt FD' iali.lL dt 

B, iolDhdrt!, rbd.l ro..ar C' D, phdk @!14, tdr.l Dd

-60. 

H..d s wid. 6 pmmtln. EyB bla.k. Promrum disrimdy wnbning in fto.t. 

L€gs ycllowbh, wilh lieht brcqnbh 1inge. Elytta rcddish-bro*n. {ith a pale hln al 

bandi donal ield provided vith 3 rie veins, lst dilided into 4 - 5 bhnchq, Cu wit 

2 - 3 bramh4, rs!t& veinl.B nun rcu5. forFing aguld dmr tqur. ells: lat 6l 

field brcwn. wnh 12 - 16 yellow lircs,lsr6 _ ? b.ing br.nches of Sc. OvipGior rcrv 

loog. alnost d lons s body. its apial valves sMll, smooth, laMolatt and acutelv 

Meslrhcnts (mn)r 4d s9 

s,D. 

23-Vlll-94, changa 

9, 

22-vlll-94. cn n8. 

r.03 25.G29.3 

,r.nro.t 

t6,23 

',;" 

s.D. 

0.33 

"*

{l- 

P..rtF6. Ahag{ (roba'rek SinSh) I 9. l9-Vt U-94i Cha.g! MmSr (r'.s!r) 

2 d 3 9, 22-vlll'94: 1 d I 9, 23-vlll 94. 

Tnc ftmb.6 of thi5 rp@i6 wrc found in t[c laf-lircr ad rcsing logs undcr 

a deNe forc6t plantstion Nnd lighr soure Mr a for6t. 

Dininglishtng choadeB: 

h h6 r very cl@ G*dblaE io t drtar6, but diffds in lnc follNing 

He.d wi6 @nspicuols yellow band sloig interulmargin of @h evci promtlm 

wi$ latd.l lob6 nosrly y.tlryish. vnh a wry blo*i band on sup.rior mdsin: 

dimr loucr l|lan wide, laEral field *nh l0 - 15 veim, lst 4 - ? being bd@llat of Sci 

wnss very loogi nale gcnitalia (F'es.l5BCD). 

Tk Mm of dit sp€i.s h4 b..n .Ltiv.d an€r $c MrD of Dr. Muhdmd 

Yousul {ho n an enirc laotomisr in Paldnan- 

Thc rypc m4cri.l hd bee. d.posit d in lhe Ins@t Museun, Depa(n"r of 

Agridhohl Erbnology. Uni*Rity ot Agriculbrc, Faidabad Pakisr.n. 

MALE: 

4. Ll@{ruut Pocq8 

(Fiss l6A_D) 

sP. noL 

sie lqrye (21.0 nn). Gcm6l body coloueion bl.c*ish-brNn. 

Na.ly s wide d pronotum, blackbh_brcwo, witlDol anv orM.n|.tFor tace 

emn€ly €ddish-blwi, wni anteltTeus yallowish; clvpefre.n! sutue stligh! E B 

pale yellowisb, febly spottcd with blrck Ocelli *hitisi latcral two alnosl rounded 

median lengthened laterally. Muillary palpi ddt brown Antenre wnh lst €goent 

bhct, rst b.own, thcir skeh yell@ish, ckcl.d with bl&k.

-a 

Pronorun nuch eidd the long, with sides very reebly b!l8ii8 out, ant dor 

ma.eii slightly @Mvq pdt rior sinuaE, both &n d eitt dee rows of brbdBi dist 

@nvcx, bl&*, dc@ly plbceni; latcEl lob6 of 0E sne @lour d |h. llrt r' but 

slighrly faint at antcrior egle, inferior mrgio ending polerioly. 

tlgs conolorou sith !ody; s€rior tibBe p.rf@l.d with a brge, oval .tl.ml 

and a sh,ll. dndcd inrnEl tyip@nr Pdt .iq r.no6 shon, $ick, siolt, fccuy lishr 

@lorcd, sniacd virh ddk brown, fi€ iomet i €rupt d by two mrcw, lonsitudinal 

stips on cxt rml eid.si pqcrior tibi& tlightly shon r th& f€noB, provid€d 

'€llowiin win E dierEt, 7 int mal tttlng spiB, which N h@k d d ot.it !pi6: .pi6l sp6 

long and stMg, superoi ermls 0.81 tirus ns long s oedioi.Ei.als, the laltcr 0 7? 

timB d loDg $ pct .ior bsitaEu, n diaxt tnals 0 44 tibB a long s tl. lllt r, 

which is previdcd with 5 inlcml, 9 eit rMl denticlB. 

Elylra exEnding to ti. apex of abdom.n, brown, witi light laleral al6i nirror 

much lonsd than wide, aldost oval, divide

It. 16. 

5l- 

t t4ttt lat tmc@ a @.l, drh 6; DD, r!a.b dj 

i'.r.*l!., da.l.r.d: c, q !LD. odd-, r.i.l rd

M.smDo|ts (m)! | d 

'64- 

r,!, tt.w. P.L. P.W, E.L, W.L, M.L M.w' l.f.L. A.r.w' H.LL 

2t 5.7 a.5 6.0 r1.2 U.2 2.9 2l t..S '.16 tl.5 

H.W/?,W. P.L/I,W' A.LJIL M.LJM.W. ll.T.L/ll.F.L 

o.ts o.?l 2,9t l,t8 0.7, 

Mia.rl.l q|ndncd: 

HohaJp.r I d, 22-vlll94, ChdSr Mdt. (Krs"r) 

A sinslc n l. of tn! w tpcc€s hB ben colld.d tun dE Lrf-lina urdd I 

Dblnrubltq.n!!rcle|ll 

ThG s.iitli. of d!i! lp.cias lh dFn cb.. finitv with d$.c of I. !m 

rrom whict tl|cy 6n @ily b. !.PDrd by O. following cotrlbinttion or chlr-E r: 

gody coldr bhckish-btM: hdd wi0ort vcuow cvc insi h.. l$g.lv rddilh- 

brcsn: Glyr.r with .pkal ticld ttry lon8, 2 93 tiB s long 3 Prenoun: po.Erior 

f.n@ 3,15 titu s lon8 s wiiL; P6r.rid ribir eitt 6 irEr, 8 dEr !tiB; &nidir 

(Fi8!. I6BCD). 

Th. n.eof thi! 3p@i6 hs bei d.littd frei! it!.pitl f.ld of .lyrn *hich 

Tic holotypc nEb h$ ben d.ro.itd in tlE ht€ct Musutn' Dclu!trcnl of 

Agrbslur.l E onology, uniEBiry of Agddl$E, Frbdrbld' Prtill|n. 

5. T.Lotrtuut lggetdtt .9' tor. 

(Fi8s l?A-D) 

MALE: 

Sir. ldg. €3.0 hm). GcF.l bodv colddtio. ltEbdiDg dB! of lc$

-65- 

Blackish'brown, wi6 a faindy viribl. yellow band alotg $. inrerial megin of 

€y*, which €xtcrds ihrough l{.rNl @€lli alorg thc si&s of ftonol rocEuni adput 

adomed with 6 vciy faint yellowLh lin*r face dark r€ddish.brdn, Pr.sntiry beneath 

rhe sterior @lhs a dim tcllow lirc; clyp+frc al sutuE sraighr, Ey6 brcwn, sFttcd 

qnhbhck. Palpibo*n, the mdilla.y with eloogate s.gmc.6, 3td lnd 4th ndlycqul, 

sth nuch lorgcr, febly wideninS. obliqocly lruMt . bl&t !t ap.x. AnEn@ brcwn, 

witb lst ahd 2nd segnen6 blek. 

Pro.oNm suugly develop.d, wid..ing in front. its ant rior mrgin fe.bly 

@ndve. pdr4rio slighdy einuL, bot tmed witl rcddbh-brcwn brisd6; di![ bl&k, 

slighlly @nv.x, prcvid.d with silky brownish-rtllo* p!b€s4@r lltct.l loba 

@n@loro6 wio lhe dhk. infedor mSii de.ding backwatd. 

Llgs unicolourour, bre*n with blackisb tinSc; ankriot ti6i& wi$ ! la.ge, ov.l 

exterMl rynpanun, *lrici is rcducEd 10 a sraller, rouoded one on int rtul fei 

poneiior femon short, srout, bleck at rpex, 3,07 lio6 s long a wide, sri.Ed eith 

MN, obliquc yellowiih lir6 on ext rrtl siddr PGt rio. tibi&. Psenting 6 i.ner. 

6 ? ourer ipin6, vhich ar€ hmted at $.ir apiclei api€l spu6 wry long, 

sup€roincrMl! 0.91 tins s lons 6 nldiointernals, *hkh te 0.E2 tin.s s lonS $ 

pd@rior b4i(iBB, supccxt rnals 0.69 rim* s long a nedioinlcrFh, ehicn tr. 

0.57 times d lorg s pdterior bditarsB; thc latrcr dned wilh 5 - 6 imer, 6 - 7 ou0cr 

Elytra shorter th.r .hdohen, rerching thc atq of eiehlh abdoriEl tergum, 

ra6luc.nt, brcwn, wi6 pale lal.El bandsr ninor snall, aln6t oEl, slithdy lone$ 

6an wide. divided at niddle by curueil veini diagoMl vcin sLahht. ft€bly cwed at 

aorcrior angle oa mirrori lsr and 3rd chords lceblt and r.glldly, while 2nd st onely 

cutued, lsr unh.d by a small vcinlei dcl) with diagonal vcid dd mirrorr harPs wnh 5 

obliqu.v€ins, firer 3 shngty sioua€, {cUi.velop.d,6t f4bly vhibl€,.lh6tsttaighti 

apiql fi.ld w.lldcvelop.d, 1.29 tiner a long 6 dc mircr. presenting 4 v.ins, 

icriculation aormed ol Mnow, clonsal!, almd r.guld cllr (Fig. l7A): la@dl6eldPalc

cwn. wirh a dat brown bdd along slpcrior olrgin, prcvided wi$ ? cle, rcgulrly_ 

3pac.! veiE, S.3-brNrched. Wing! rery !mll. shorter than.lytr. 

D&k brcwn above, yeuowish-brown b.neath. Csci dark browi, very long, 

I -6 rim6 s long d elylra. Ccnitdlia (Figs. ITACD) sinile to th. p@.rling +@ia. blt 

difllr in havins thc p6Erior median pojction shod, b@d .nd neo*€d abtuPdv 

fonnin8 an ohuse n dian point. 

Me$u.denls (m)r 1 d 

B.L H.W. P,L, P.W. tr.|-. M.L. M.W H.F.L. tl.tw. H.T.L 

6.0 lz.5 27 25 lr'3 a5 los 

H.wJf.W. P.L./P,W. E,L,/P,!. M.L/{t{ W. H T L/H rL 

0.95 077 2.72 1.03 0.16 

tloldype I d. 2t Vlll-g4, Lrl Sol'dF (B.[awlpur). 

A singl€ nalc of this sPei6 bd b@n coll@Ed trom thc lcaflirl4 under a d€ff' 

fore$ pra.tation of sufaida (Ercqltptlt cawldaldsis) 

Dletingulsbing ch.roct.B: 

h sh@r i|5 clc. l!@blare b T.ko|rylrs pN.N sP. @ . fiom which 

ir diffeB in lhe following ch3@B: 

Head pres.nling a fably yellN band alon8 thc int€rml margin of eves: ccipur 

]!irh 6 sholi, f@bly visibleyellow linesr elyrasho €r than abdoheo,2 72 tim4 d long 

as p.onotuni nirct I.08 times 6 lons a! wide apial field compdairclv sho , 

0.28 lire s long 6 clytra: €piphallB wi$ shon ad widc m.dian p6Lrio. psjdrion 

lFig. l78). 

The name ofthis sp@ies hs ben dcrived tios th. faintlv vEibl. vellow lin$ 

*[ich are present on th. occiput and along $. inErnalMgin of.vs

I,r& 17, 

47- 

f.lotAgtt WC&4 4,r.A,drlt a; &D, rrrd. d; 

x, dtLr+ dd.l.{.d; C, D, ,bl!. odcld, t dd .d

{E- 

Th€ holoryp. nrL of Lhh specics hd ben deporiEd in the ltr4i Mueur. 

Depdlme.t of Agricultu6l Enbmology. Univ.rny ol Agncul$re, FsLalabad. PaIsah 

GflN ftdts"/llr Cbop.'d 

t r. Plate,rllrt Chop{d, EG, M!d...3?:2?5. 

'trlEt GrJIhu qudnnisatw SN9rt. 

Aftording to Ot! and Cade (1984^\, Grylld lineaUc.pa Walkd, ued s $e 

ryp. of this genus by ChoPdd (1961 , 1967) and Rud€ll ( 1964), i! a prinury homonvm 

ot G. liieatk.ps Sral, So, it snout bc rcplacld by th. onlv availrblc svmn n ol 

C. qu4dtisdsata urdd this senu. 

This genus h rcPlee cd by 3 epeics, which gtotlv hllv wilh.he publish.d 

dcscriprion ot oltc and Cade (1984a) excefl $e Iatedl ocelli, which are rd€lv @nn4i'd 

wnb rhe yellow sldpe, f@ snerims yellowbh, elytra cxt nding up to fte tip of 

.bdomcn i. both cxd and slidt difrdng in l|BftnoE, 

L 

K€y to rh€ slqies ot I'l4ttyfuls Chop,dd 

body @louorion shrnrng black: he$ *ilhout an, mdkrng 

..,... h.ladoc.phafut (SedllL) 

CeEral body @lNltid brewn d bL.IBh+own; h.!d with b@d 

doBun d shon li6 on @iput. 

z, Body sonewhat deprcsedr head with 6 shon, longiodinal, vellow lin6 on 

eciplq clypo4ontal suture strongly ugulaEi ovip6itor sbon, with aPrc{l 

vall6 bh.t.t.pd (Fis. l8D) Dad.,t (S.us6an) 

Body clong.Ei hed with a dow, tr.Nwe, ycllow bAln bc$q cUi .nd 

a broad brnd on v.rEr; cllTeqftontal sulure feblv a.gulale; ovipGilor verv 

'ong, wirh lpical valveq naiios, acurcly poinEd (trg. 19D\.

-69- 

L naq,r'rn(s D.bnoc.rh.t6 l*.aflcl 

1838. Aryus rula@c.ptun^ SeNillc i@ salsurc, lE??) lc. O n., P. 342 

196l- PlatygrJllut n bn6.ph4l8 Cbog&d, E6, Madr',3?:275, 

Thc oily fcmale colla€d sp@iftD gddy taUis wilh lhc publish.d dscription 

(Chopard, 1969) of thir spcciB cxept thc lae and marillarv palPi, which arc brcwnish- 

blek, elytr. d long ar O. abdomcn in fcMle atd Sc 3 _ /t braMhed. 

Mesuremntt (mm): I 9 

B.L. H,W. P-t- P.W. E,L. g,r,L H.D.W. d.T.L OL 

3.lo 2..r 4 30 9 0 9.0 6.10 9.50 

H.W./P.W. P.L./P,W. E.L,/P,L, H.T.L./H,! L. E.r'L /O.L 

0.63 0.60 3.10 0.71 0x 

tsldabad 19. I l-lx 94, 

A sirglc f.&le of this speiq *6 coll4Fd fiom gra undr a light eurc!- 

This spdia3 h4 ben @dcd rd tlE fiRl tiD ftod P.litra.. 

Chopdd (f967) shified dis sp.ci6 frcn C'y4t{t 6 PldtsttUus, b$OntarJ 

Cade {1984a) mcnrio..d n a a doubdul mcnbcr or $is &nu 

2, PlttrmllB ,,ar&n (S.@) 

(Fit! l8A'D) 

t817. AryIs bwi Si6uG, M.n- Se. G€eve, 25: l?0. 

lgtl, Achad btuice.la fwny, SB, Atad, r,/ti!s. wien. ll6i 288. 

l9O7- Ach.ta Uwti Kdny. lden. lbid.. 116:289. 

1943- CryUul6 ,nrt.t Cho?&d, Orth, Aft. N., P. 190 

1964- Pl4ttgt U6 ,@ti R&xlel, c{t d. Ent., 95(12): 159E. 

The @llecEd material largely confoms to the Publisbed descdPtion (ChoPud,

hl!!xFl|d!d4& 

rt. rE 

.?G 

g#,ffi#3#iilffi 

D

1969r sac.d, 1989) or Ot speci6 

0,83 - 0.89 tim€s as long a widc, 

blafth6. Gcnitalia (Fi8s. I6ABC 

M.su@enl'tm):5d 59 

). 

-? l. 

ehich hart 4 - 6 htrP3, ni.lor 

5 - 6 venB and Sc b.ring 2 - 4 

s.D. s,D. 

,.nro., 

Khuzdd 2 9, ?Jx-93i Kohsl I d 2 9, 3-vlll-93i Pshswe I 9. l_vu-93; 

7 d 3 9, 2-Vllt-93: TarMb (Pshawat 2 d 1 9, lTlx-9ri D.ta Isdil Kha. I ?, 

26-lx-94: Bhagar CToba Tek Sinsh) 3 d 2 9, l9'VUl.94; Changa Matga (Kdut I l, 

l5'Vl93:5 d,22-Vlll-94i ! d 5 9.23-VII!-94: Faisalabad t 9, l4_Vll_86: I 9, 

l6-vll88i 5 6, ?lv-95r Cut*ala (Faisalabad) I d I 9,2l"vl_93i LahoE I I,

72 

25-vl-89r Lal Sohaora (B.hawalpuo 2 d 4 9, 2t-vlll94; Layyah I 9, 25-vl-89i 

P.kpauan I d, l0-VIl-88: Rawalpindi 19.5-Vlll-88i 3 d 15 9,7-V|U-8E:2 d, 

8-vll!E8: I 9, g'vlll88; I d I I, l5-vltlES: I d. 20-vlu-88i I !, 23 Vlll-8Ei 

| 9, 25-VIll-8Ei Sialkot I 9, 25-VI-89. 

It is th. mct abu.dlnt sp@iB found in dlc l@fint.r utdd deN fot6r 

pranhrioN of shahm (Ddlb.ryia tit@), !{4t{ eopr!6 rpp.), bdb@, 

lDatd@atdis stictit),s!f^\d^ lErco\ptw conaurl%k). ett lt wa aho @ll4rcd 

along r@d sid6. in h.dg.s lnd on grues d.t light su@s. 

All rhe above ldalitiB .xc.pt Fakalabad, Palpattai, RawalPindi atd Sialkot, 

Chopdd (1967) shiti.d dn spei6 fton Cryl&r to Pl,tteryllla, b!.O@.nd 

cadc (!984a) ncntioned it a! 'Nomim Dubia'dre m ftc non{vailabilily ofde nal. 

type sp€cinen. The lp@ioers coll@ted from this rcgion difiet tom lh€ Publishcd 

dNriprion (chopdd, l%9) in a few ch.mt B. bur a fie mal! gcnitalia detly 

co.lb.n ro lne publishcd fieurc oa ihii sp!.i6, $ey dc plaed undd $e @e 3Fci.s. 

MALE: 

3, ndts.tAus blconpaa sp. not. 

(F'g!. lgA-C) 

Sire nediun (16.5 nm), Body elongal!, with brown @lolEtDn. 

gl@k, prolid.d wih two ra6vers. ycllow bands, oe b.twen latcral @lli and 

the other on vcrtcx betq@n lhc upper m&gin of €yes (Fig. l9A)r t@ yellow, qith a 

rdisv€e, bown paEh bcl* the clr?@frenal sunte. which k feblt @nvd, the 

lau* @lo!r.xt nding er.riorly in the fo.m ofa lic up to the scdian @llu$ fiont l 

rcsiruin blackish-brown; ch.ek aM bandiblB yellow, EyB pi.kish-ycllow. Marillary 

palpi long. yellNish-brcwn, wnh 5rh stm bl&kish-bNn, obliqlely trundoe at 

aFx, Ante.@ bown. witn l$ *gm.nt Fltw. febly monlcd with b@n.

-73- 

Prcmtun tr.dv.se, 0.?0 tincs 6 loig a w'de, sligldy wlbning in iionl, 

anErid mdgi! vdy febl, @@ve. port fior dndt straighl; disk nat honled wiin 

y.llry ad brow., Ery iEly pubcsnt d gla!ru; lailr.l lot s *irh uppd half 

brown, loftr y.llow, €xtrere iiferior maBin black, febly ending poatcriorly. 

L,gs y.llow, d.Nly pubsent. 8pou.d wnh brcwni dnt iior tibis. amed wi0l 

a snrall, @unded inncr and a lug., oval out r lymtdnudi antcrior and niddle hrsi 

provid€d wi$ rwo rows of stull brnds on v.rEal sidq pdt rior lcmda addned *ith 

obliqle bw; !t i& on both sid6, incmpt d qid lwo lonsitudind y.llow liH on 

cil. nal surf@i po6rdie tibie.nEd with 5 im., 7 8 old spi6, ncdid eical 

spur of bod sids lorScr thd oth.rs; polErior baitarsi0,4? riEs a long a tibiar. 

Elytra l' lobg a abdohen, li8ht bowr, aln6t u_aNpd. ; mirror ndly @al, 

much wider rhan lo.g, divided slighdy bcloq middle by a curyed veini diagonal vei. 

!e.y lo.gi chor.ls cguldly spaed. curvcd, lst metitrg $e dago.sl v.in by a stull 

v.inl€q h.rps with 4 febl' sinut 06liquc vciB;,pidl fald anhcr lMU, provid€d vilh 

o.ly 3 veiN, ..ricul|rion foded of nrcglld, oide @lls (Fig. l9B); lncrd lEld Pal., 

eirh upp.' dargin brown, prcvided wirh 6 - 7l@blyap@d, rcsuld rciB, Sc b4ii8 

3 branches, Win$ vcry lon8, @!dlt., 

B@wn dov., tcllo*ish b.Mrh, Cqci blek. Cenialia (Figs. I9EFG), epiphallus 

rilobr! apcr, dEdd bor&. dccply lnd nllrwly eMgi@l!, wi$ ultriu dN 

cuN.d upd&, p6t riq @Ci! wi! la|.nl lobcs 3horr, rn ftdiat &rslly @wcd: 

@logaraGGs short, dilided i o two lobca ait riorly, hidd€! b.ne3o a'phalB in 

FEMALE: 

Il exatly t llis witlt ihe nalc .rept ttc follosing ch,rei.B: 

P.onoru.t 0-70 - 0.?4 tiB s long s wide, its lat ral lob6 lrrgcly blwd, with 

a narcw y.llw bsd at iiferior Dargin. Poskid tibia. prwid.d wnn 5 - 6 inrcr, 

? outer spind, Elyua s long s or h{dly longcr thd abdon n; dotel ileld providcd

Fig. 19. 

Plattg.lAut bi@nptas sp. w. î rc d; B, dnn d; c. .h. 

!.4 e; D, .npodror trtt-r E4, rolrtl| dr E. +!tdtE, 

ird Grd; P, G, lrdrk 6ndq rdnl .nd t !-d *< 

c

-75- 

wirh 4 fr* EiN, Cu 5-brarFb.d, Eguldly obliquc dd c{uatly dbonr, tr$!vcr!. 

ltinlcB nun€rour, fnnilg in g!ld, .lo.g . elb; ldcnl ficld l,ith 6 EnB, Sc 

s-bflehcd. Ovipo8ito. sy lon8, .uiight, iB apbl wlver long, ffiow, &ur.ly 

poi .d (Fi!. l9D). 

M.dttto.!.s (m): ld 39 

Md. 

l d, 2l-utr-94, 

I 9, 2l-Vltr,94, 

2 9 , 2tVnI-94, 

|-t2 

,,-*rr,o 

92-tO.2 

t3,0 

3,5t 

,1,, 

t1.31 

3,30 

Ld Sol|,rnF G.taedpu)- 

L€l Soham (B.hawalpur). 

ld Soh|ra (B.lbulpd). 

Thc sFcih.d of rl ! rp@i!. *@ @llal.d liln dE lc.fl'0.r unda d.|!. 

sD. 

0.t2 

o.31 

";

-16- 

foBI pldladd of suf.ida lE|uqpt6 .arsAub8i:, td shehh (D.lr..sria 

DbdrSubtirg chrrad€E: 

flr goiLlia havca cl@ rd&nbtare to rh@ of Plalttryttd b@n,btnn 

dif€B signifi@ndy in th. folloping body cb@r.rs: 

Body .longarci had sith two t dBw.s. y.llry bdki cl!?.o.fionul sururc 

sliShtly angulate; mnror dislincrly widd than lonS; h.rys wirh 4 obliquc vciBr 6piql 

ficld with 3 wins; late.al {icld eirl 6 ,7 wirs: ovipcior very lorg, irs api.al rdvB 

Thc Mc of thir lpabs h6 ben deriv.d fton Oe prc!.@ of tso r.sdsc 

Th. rt?. naocrial ha b@n d.Fsircd in rhc Inser Musun, D@dtmcnr of 

Agriallunl E tonoloSy, UaiveBity of AgriculiuE, Faielab.d, Parbun, 

CaB Ph.Uolobatut stu @a. 

TJP.I Phat obbosE ad.ntibialis 9.n ... Np. N. 

This gerus , *hich ir rep@nl.d in Pakisld by rwo mv sFci.!, is.rccEd oo 

rhc bdb of ndc sedr.lia. tt tcys our in R.!d.U (t954) ben"ea ptatsryUb sd 

A@nthotryllB, but diff.6 fion, the forner in endopramcr.s, which e withour 

mNvce panncEl Eu$lc apodcG, rrd rhc laiEr in msal lob6, which h.vc rh.i. 

.pies betw@n the apies of el.9&do6. Thc c,piphall$ show! il! clsc affinny b lhat 

of Achao, blr ir @n 6ily be ep&ard fon rhc latir g.tru by rhe folowing 

Epipbatlus diltiDcrly vidcning p6r.riorly, ati€lly b.arins fivc tobd (orc 

b.twen thc mcdiao ard hr€nl lob6 or @h sid. and v.ry fe!!y liriblc), basat ponion 

prcduccd upwads giviig it a dc.?ly cup{haped appwa@; inrcrtul anr.rior apodems

-71- 

ve.y longr 4lotemG vcry snall, wid inEml .nErior pro!.lioN v.ry long, thc 

lauer oriljMting nar thc ba6 of th. formcci nesal lohs wirh then spic4 bdwen dc 

api6 or $e et p@mcc: eldopa6re6 C-shrpc! (Figr. 20BCD). 

Othcr diagndtic fcaruB de Body @lou. Eddkh-bben or broM; hcad with 

or without a yellow band bctsEn @lli or ycllow lia oi @iputi premtln 1.43 - 

1.67 limcs d yid. a long, ils lnerior margin diltircdy coMv., posterior alnost 

slniShti .lytra r.:chir8 to the ap.x of abdond, oirro. alwaF widd rhan lory. dividcd 

by a single veinr wings lcry lonS; adorior libiac FrforaEd on bolh sidesi posrerior 

tibi& dd.d wilh 6 inq, 7 @l.r spi6. 

The me of lhis g.tru h.3 ben ddiv.d ftoh ihe .piphallu. which i! provitl.d 

wid 5 !ob6 o. p@rc.ior margin, 

Key to the spe.i6 oa PhaUolobosut gen, Nr, 

l. Hcad wftout traBErs ban

-7E- 

browi, mottled wi$ black. t t ral @elli rounded, ycllowish, th. antEdot lllltauy 

l.ngthercd, Mdillary p.lpi bwi, 4th s.grc sho q rlu 3rd and 5dr. O. l.tEr 

feebly widcning a.d truncnte at aper, A enMe b@n, with firsl two slgdenb reddish- 

Pronorum wide! than long, dislioctly w'deoing in front, ns anterior margio 

ltronSly @n6vc, pGr.rior str.i8ht, lne lan4 lin d with )tllwish: diik almd flar, dek 

bro*n, *ith reddish-brcwn pyriforn impBions, and ! yellow 3pot nd p6tctGlaldal 

atrgl€s: lat ral lobd ldg.ly ddk brown, wi6 a @rcw y.llow linc along inferior Mgin 

of th€ antcdd angle, tlt? larl. MSin &nding pct riorly. 

kgs of th. sne colou a! $e bodyi anlcrior libia. with . rynPanuE on inner 

aid otrt.r i,6, inier one smaller, roundcd. olllt onc very h8c. elongat r PosErior 

legs rarlEr shoa, Lmora thick. rtrong,2,46 tim66lone s wnk, srialed with fcblv 

vhible ycllow lines oo eiterMl facs, more promiftnt rcar ba. add alonS lower nargini 

posl. iq rib@ r.latieely sho(, 0.66 dnas 6 long a! posErior fcnoB, povidcd *ith 

6 inner, 7 oul€r strong spiB, slp€toinrcrnals al6Gt eq@lto ncdioiitcmab, which arc 

0.8 tims d loDg 6 posl€rior bdnatsus, dE land prcsenting 6 - ? inner, 8 ouGr smau 

Elytra extending lo thc tip oiabdomen, brownbh_y€llow, spon€d wilh brcwni 

mnror nuch widcr th& lo.g, it! ..gl€s almdt round€d eic€pt $E oucr oft, divid€d a 

lifite behind niddtc by r cw.d v.itr (Fig. 20A): di.goMl vein rct ltaiglt. iliShdy bcnt 

lo*ards ln chordr chor& regululy curved, lst alnolt touching thc mi.lor near ioner 

mglei hlpr wilh 4 obliq@, linuaB vci6i ,pical ficld weltievcloP€d, pr*stinS 5 

veis. reticulaion formed of almct pe lgon l. ircgllar ellsi laEral t.ld bl4kisb- 

brown, provid€d with 7 8 fably and rcgulely spac€d veiN, Sc 2 ' 3 br@hed wings 

greyish-whirc. vcry lor8. l.?l rim4 4lor8 d clyra 

Bhknh-brown abovr'. ycllowish-brown bcneith. Ceici dek eddish-brown. verv 

loog, O.E2 tims s long s poicrior fenon. Genillli. s dc$riu undd 8cnB. wilh 

epiphallu having sidcs sraighr (Figs.zoBCD).

t.2!. Pr,/&J,dotut tdaultttb !4. . t. d. & ittr dt 

tD, !rr.n dr 3,.dotI!, ac-l .a.ct C, D, ,Lrc 

d.eLr, ndd rd llr.l .t d.

M€surcment! (m): I 6 

EG 

P.W. E.L. rV,L, M.L, M,W, Ii,F,L, H.F,W, H,T,L, 

16.0 5.0 3.0 5.0 r0.5 13.0 3,7 6.0 

II.WJP.W. P.LJP.W. E.LJIL M.L./M.W, II.T.Ltr'X.F.L 

t.0 060 1.50 0.35 0.6 

HoloitDe I d, l4-Vll-88, Faislabad. 

The only m.lc h6 hc.n @ller.d frcm light n@ thc culrivared f'cld!. 

Distinglbhing cha4cres. 

h ldb iimild to Ptot!8ryUB btuM.n, bnr 4ifid3 in dE followiig chdar.B: 

Bodt reddbh-brown, with yellorish-brcwn elyn; ptum$m widcniog in front. 

wid anrc.ior rusin rt ongly @Mvc, pGfiior $Eign! mind l.l7 rimB a wide a 

longt 4 harp vci6i apical fi€ld vi$ 5 veinsi gcnilalia (Figr. 20BCD). 

This specid ha! b@D named afier thc tosLtior tibie, which de very shon- 

The holotr?e male har ben dep6ilcd in il'e lnsr Musum, DcPdlncnt of 

Ag.icuhural Entonology, UniBity of AgriculluE, Fabalabd, Pakb|.r, 

2. PhoUolobotus ehwe8b sp. tor, 

(Figs.2lA-C) 

MALE: 

Sit n€diud (l?.5 mn). Gcrchl @loontion bwn, but lcgs f€lbly rinscd wiri 

ShininS bl&lq witt lwo i.aNv* yeuow baid!, one oa.row and on fronral 

rcsEum, dt ndinS rhough latml @lli up to.yes, orh.r wid.. fccbty vbibl.ons.cx 

bcNen opFt mdgin of cta. lhis cx&nding b.hind @h ey. rhrougn chak up ro 

mandiblesi f@ yellowbh, wiO posrlypeus brownarniddl€i clyp.o-t@nr.ls!r!rc fcbty

t.2l. Ptulblob$r nt M.nrb !o. .. rp. e, A, d}rn 3; 

a c, -rrn. a, 4 dlLla darl .r.iq C, ,hl!. .qla,

-82- 

angulaEt forlal rostrun brown berwen .nEnna.. Eya blat- Anterio. eltu! y.ltow, 

witn { small vcnic.l line of lhe sln colour bcMrh n, joining a *id. Eans@ ycttry 

b&d abovc clyp+frcntal sutur.. Palpi brcwn, with 3td and 4rh @illary 3cgne . 

nearly equal, 5th clongrtc, aebly vidcning, obliq&ly mul. and bl&k ar aFr. 

Anlcn@ bbwn, their s.ckea ringcd wirh rhe enc @lour- 

Prorotun wnbr dian long, its 3ide mildly bulgiit our niddl., anr.rid @gin 

concave, pdterior suaiShti dbk sligh y @nvcx. very pubMcnt, daik brcwn. f€bly 

motdcd wi(h glowbhi l.LBl lota wilh iupdhr p.n drrk brcwn, irfcdd tcllow, l|lc 

lalcr @lour wid.ning et riorly, .ntelior argl.s rounded, inferior mgin sliehtly 

t gs @n@lorous, pub*ent; amcrior ribi& with .xtefinl rymparuh lsg€, oval, 

inr.rml oN vcry rmll, rcudcdi pGGrior fcnora clon$r., 3.5? ric s torS s wid., 

sfiiated wirh i,row yellowish lines, iot ;up€d by a longitldiMl yellow li in lowcr 

nalf on cndml surface: pqlerior ribia sned wirh 6 idernal, 7 exErnr! rriEs. 

Elyfd r@hiDg up !o thc b6s of @rci, uulus@or, brown. with vcry nat 

venationt minor much widc. rban long, aDrcrior and posrerior !ngl6 atmt .oud.d, 

dividcd b.hind middlc by r cw.d veir (Fig.2lA); dilgoial v.in loDg, be.t.r anrcrior 

an8l. of nircrt chords f..bly dd egul.rly cuded, Irr unird eirb di.gonit rci! 6y a 

smll veiolet and rcnol. fron irrcr anglc of mn.ori harps pr6eding 3 disdncr. ore ot 

rwo obliter.t v€iDs; api€l 6eld prcvid.d wnh 3 veiN, fortud of l&g.. clorlar. eltsi 

lai€61 fjcld tra6lu*n1. yellowiih,brown pracndry 6 rcgubrlyipa..d v.iE, Sc 4- 

bdnched, Wings nuch long€r than €lytra, 1.55 rin€s s long 6 rhe latter. 

Tergites biown, mor€ or I6s spon d *ith ycllowhn, !rcrnioc! yeltow, wi$ 

b.wnish ting.. C.ici long. d&k rcddisn-brwn, | , | 2 nn6 a ton8 d po3cric fcmo6, 

Ceniulia sinild b rhe t?€ of rhe 8cnus except epiphallusi which is eirh febly 

@mpes.d sidd (Fics. 2 IBC). 

Msu|@6r.lM): ld 

M,L, M,W. B,r'.L. |l.&W. Hr,L, 

2.t 3.0 to.1 r.0 9.0

-83. 

H,W,/P.TY. P.LJP.W. E.L/P,L, M.LJM.W. E-T.LJII.F.L 

033 0.?0 3.67 

Holottpe I d, 2l-Vlll-94, Lal SohaM (Bahasdpuo, 

A siiSlc n'.le of rhis sFct s ha h€n @llctcd fon tn b.f-lirt.r ud.r a d€e 

foiest prantltion of sufaida (t a.LrptB comArt^k). 

Distinauishiog chulcae6: 

The g.nihlia of tht rFci$ show rh.ir close afiiniry with lh. tt?. ol $. Ccnus. 

bur differ ir head markibgs. clyp.qlioDlal sutue, which h febly anguhte, bulgitrg 

promtun al hiddle, .longaE pct rior le8!. yci6 of api@l fi.Id. its rericulnion.nd 

The Mrc of thi! speis ha ben dcrivld fion tft. ttP l@lily, Ld Soham, 

hrving a fordt plaotation and 30 kn aMy from tn f.dous cily of Bahrw.lpur. 

The hololype nalc of drk speis h6 b€ei &posited ia fic lnst Mueum, 

Depadnent of Agricultlhl Enlomolo8y, U niv€trily of Agricrlrue, Faislabad, Pakiskn. 

G.trG a.r.ro rlbftlB 

l?75. .{./r.,a Falricius, Syet. Enr, ed. l0,l:2?9. 

1906. Cryl6 Kirby, Symn. Cat. Onh.,2: 27. 

1935. C.rl&r Uvdov, Ann- M.g. Mt. Hhr., (10)16,320- 

Type: Gryl6 (A.h4o) doneiica Li.na.!r 

Thc @llccted ncmbss of 2 spei6 dnform to thc publish.d dcgiPriot (walkc.. 

1869i Ra|]d.ll, 1964i Otte add Alexander, 1983) oftis genus. 

Key to tbe spei6 of ,4.r.rd F.bri.iB 

t. Prorctal dark mdkings neither extending to lh. anterior Bargid nor 

otctolak.ally (Fis- 224)i .lyra eith ari@l fcld shdl (Fig. 228) (l6s lan 1.2 

rin6 a lo.g d th. mircri epiph.llus i. do*al dp€ct roughly quadrare. . . .

-6/L 

PMoral drk Mkine! cn ndin8 io tlE dt !io. dgin ald dimlatdlly 

(Fis. 234); elytn with apicsl fEld long (Fig. 23C) (mor. thin 1.3 tibe3 a! long 

a$e mKor): tPphallB in doMldprr dirnrcrly tilobrtc. 

r. A.h.t b^.ttt$ (rjr,',rul 

lFist.224-E) 

1758. GryUa (Achaa) domrt.a! Li.@B, Syst. Nar. ed. l0,l:428. 

1161, Gttlla dowttic6 Lin@us, Fauna Su@. €d. 2, p- 236, 

1175. Ach.ta dotustiM Falriciu, Sy!r, EnL ed. l0,l:280. 

r87r- Gryll4t tt@.6olit walld, C{L Ddmrpr Sal6t. B.it. Mu., suppl. 5, p. 6. 

1y35. Gry ll8 dar6ri.6 Uaov, Ann, Mag. mt- Hbt-, {10)16:320. 

Tne olleEd speines cntiely asr.. with the publBh.d dsripdon (Abdulld', 

1955t Chopsd, 1969i Sieed, 1989) ofthB specis eftpt thc n.duFmcnB of v&ioug 

body prrs, Cenitalia (Fi8s. 22CDE)- 

M.6||l!H.s (m)r 5d 39 

5,D. 

o.21 

,";,,., 

3.3{' 

s,D. 

";

Ir8.22. 

'E5- 

A.te tua$dntt A r-d -d FldE a, d.-t s; 

a, drtn d; C.E, ireb dr C, .etDbe., drnt .rdj 

D. E, pedk oprd, v-rrl .d lsdrl ro.dr.

H,F,LJO.L, 

-86- 

Fahalabad I d 1 9, l2-vl'EE: I d, ?lv-95t Kharewal 2 d, 18-vl!88i 

Palram I I, l5-Vl-E6i 3 d I I, ul.b.lled, hrci fiod siud.tra colletion h rh. 

Dcpann.nt of Agricul$nr Enionology, UnjrcBity of AgriculruE, faisalabdd, Fati.tan, 

ft is n6dy found in hu'm dwcllings ard @ly in 8By hwB, on lid6 ard 

It b @nsidcr.d to h. a co'non, widely dboibuEd .id *rious pet of m.ny 

crops (Ahmad ed Ullah, 1949i Kha!, 1949; Yunus ,r al, 1980; S.ced, 1989) in 

Paf,istan, but the ethor hd @lleEd it only fton a f.w ld.litiB ad huntr babitrtion- 

Saeed (1969) placed 124 specimens undd lhis sp.cis, but th. dbs@tion of male 

geniklia in ihe pqcnt 3hrdi6 rdal.d rhat only 5 ip@ihe6 rc€ &hally of 

A. dowtti$ abd rh. rc6 i.ing probably h.long.d to lne ctosely el.r.d rcw sFcks 

(4. thot@ico). h n tnthq i^l! 6tins !o note that Abdluah (1955) lid Saced (19E9) 

@llded thc lau.. Mdy fioD ligha i! dr cultivatd fEldr ard tn. F*nr aund, ioo, 

@llcted i large populatioo of diis rew rFci6 fron the 6clds, It rnows $at rhi3 mw 

spei.s is rcurny a pBt of cllrivared crogr and duc !o irr v.ry cl@ rBcmblane it is 

niside ifad 4 ,,1. dru"rtirG by rhc ptcvioB wdtc^. Th! ida is funnar supDorud 

by Chopard (1969), who rcmadcd $a1ir iE rath€r lurprbing rhat tne htl. spei.s hd 

b@n @ordcd a a p6t of cropr in Parbh ( hmad and Ulla!, l9l9). 

2. Ach.u thoturi.a sD, no\, 

(Figs, 234 F) 

MALE: 

siz n€dium ( 15.4 - 2l.E mm). Body rcmcwhar depEled dd brovni5h-y.llow. 

0.t2

-87- 

Bl&kilh-brown, 0.95 - 1.04 tin€s a! bt@d d prcnotur!, with a Miiow ycll 

band bdwcei ledl @lli, anoth* bed om on d. venex, dt nding behind thc cts 

up lo chdbr f@ dst bmenr clrT.o-frontal suturc cuncd (Fig. 238). Eyca darL. 

Mdilldy palpi y.llwish, qith ldt sgnent duk md and obliqucly ruMlc at aPex. 

Pbnorln r.asv.rsc, 0.51 - 0.@ tim d long s broad, iB dl.rior 'rutgin 

slighrly concavc, pos€rio. alm6t sEaighrr disk n 2Jly fla1, with two laige dart brdn 

spors on pyrilorn impE$ioB, .xLDding att riorly and anterolatcrally, alm6t0oucltiq 

lhe dErior mngin, amrhcr small brewtr spor b.hind @h ioprcssion (Fig. 234)r hE6l 

lobs yellop, with a b.o.d b@n band i! sup€rior pan. 

Legs yellow, cove.cd pith ddk pubesen@r aftrior tibiae witt ! larg., oval 

exdMl, and a small, rolndcd in&lnal tyFpaoumr pdtc.ior tibiae arned with 5 _ ? 

inErnal, 6 - 8 qllrn l spin6. superointerMl tpiel spuu d long 4 nEdioint rn L; 

p6terio. bditaBi long (4 mn), enh 6 inl.hal, E eie@ld.nticls. 

Elytra 4.12'4,?E tih* as long s pbnotum, extnding to the apex of abdoneni 

mnror 0,91 - l.15 imes s long 4 br@d, dividcd alm$ in niddle bv a curverl vein 

{Fig. 23C)r hdps with 5 - 6 obliqu. v.isi lst chord unit d with dugoial vcin bv 3 

snall veinleBi apicd fi€ld rcry lotrg, 1.35 - 1,58 riB d rone 6 nmor, p@ntmg 

4 - 5 veiN, wi$ elongat€, rcgular r€ticulatiorr lat ral 6eld 9alc, alm6t tJadpar.nt wih 

broqn band rloig supeior lnafgii, provid.d with 5 ' 6 veiE, Sc 3 - 4 baichcd WinSs 

long, mEn bnga ths.lytra, 

Broqn above, yellowish ber41h- Cdci long, yellowish, verv hairy. Subg€niial 

plar slightly conpressd nar tipi epiphauus ttilobed, i$ n.dian toleriq poj@tion 

lons, bi@d at bse. ab.oPtly u.owing ne Pd€ior Mgin and rorning rn &ucrv 

poinred. dorslly curv.d apcxi laEral poj@tio6 slightly 3ho.t r than lhc m€dian 

one (Fis.s. 23DEF ).

tu. 23. 

-E6- 

O-AO 

A.h.ta tho@ha .D. noy, A. bad dd D.wrd a' d.r..l 

sp.dj B.lG dr C, .lyh d; Dr, 8Efrdi d; q.olle.n!' 

d..sl 4..4 E, F, phdn. ond4 Edrr .'d l|hl .rt dr'

T'EMALE: 

-89' 

Sinihr io Dlc exc. in! fouwing ctEn GB: 

Elyta wi$ Egulely tpa.cd, obliquc vciB. {irh 3 fte, rhc 

Cu 5-bB@h.d, tnnrwe reinlen nun rG, fmiig wid., 

ovipo6iio. lon8, snrigh! wirh tMc@taE.piel vrtit.. 

Ms||tears (br)r 5d 59 

f,'.L 

t2.y 

t9.1l,.8 

,.nro,o ,.* 

H.ktyp.: I d, l9-vt-93, Fairalabad. 

Alottpe. I 9, lM-93, F.lrlrD€d, 

Puttp€s: Lab.h2 d,9-lX 93; Zhob I d 3 9, cx-93; Sibi r d r 9.7,D(,93j 

r\nuzde I d, &IX 93; Abbo&bad ! d t 9, t+D(-g; (oh.r I d, IGD(.94; ziet I d 3 

e, l?-lx-94; Bsherlpur I 9, l5,V-89; Fabrtsb.d 3 d, l4vn-88;20 d 2i Q, tcvrr-88; 

s.D. 

2,t5

-9G 

I d. I7-vll-EEi3 9,29-Vlr88r I d, l7-v 92i 9 d 19 !, l9-vl-93; Gojla (Ioba T.k 

Sincn) 2 9. 28JX-EE: Kh@ul 3 6 20 9, lE-vll-EEi 2 9, llvll-EE; tdrorc 2 !, 

2-VI88rtyyahIdIt, l5-VI-89i Mianw.li Id3 9,4-X-88; Mitha Tawm 

(Kh6h.b) I I, l+X-94;2d, IGX 94; Mulai I I, lt-V-9; Mune I 9. l3-Vll-93; 

Muafiargrh I d 23 9, 20-vll88; Pakpaun 2 9, l5-VI88r I 9, 29-vl-88; I d. 

8-Vll-6E: 2 9, }Vll 88i I d, lcvl-6E:2 9, l5-vll-88r I 9. l5-V-E9i I 9,29,V1,89; 

Rajanpu I d, 22-Vll-E8i Hed Suldn.nli (B.n.walM8&) I 9, l4-VI,93i 4 d, 

unlabellcd, taken froo snld6l5' @Uc.ioi ii 0E Depanttrc of AgriculrDal 

Enomology, Univesity of Agricultur€, Fakal.bad. Pakirta., 

A very common and widcly disriburd sp.ciqs, nosrly collccrld on liSh mar rhe 

cultiv.t.d fiel& of whar, oil s.rds, v€gellble3 .rd also und€r ga6, lafliller sd 

Dislingnbhbg cbmcc6: 

ft h6 a cl* r*nbla@ b ,4. d,u6ria6, iiom which it dife.s mainty itr th€ 

@nfigrarioo of ml€ S.nitalia (Fiss. 23DEf), pronohl mkinss (Fis. 23A) ard 

!elativ.ly long apiel Il.ld. 

s@d (1989)pl@d many specinens of $is sp@i6 under ,1. do@$icu, onthe 

b6is of rheir clGe body rsnbtaD ro de larrcr, bur the aMtysis of thcn g.niblia 

reval€d 6 thcy &tully belong.d b rhh re* spei€s. 

Tle sp.cine6 of this new spaia @UeEd in or n4 $c cutnvaEd fEld and ot 

fie thr.shing fl@6 by th. pr.viols worteB rnd rhe p6cnt author ni8hr bc the p.sl of 

a8ricultural rcps dd w.r€ Dlidentified s ,,1. doroticB, wlicn is , *orld-wide 

TIc Mmc of lhi! speics hr h*n dcrivcd fron iE prcmtll MkirSs, which m 

quite diffcrcnt from thos. of ,.1. doarri.6, 

Tbe typc malerisl ha ben dcposircd in the lngt Mu*um, Depann nr of 

ASricultual EnonoloSy, Univdsiry of Ag.iculruE, Faielabad. p.kir6n,

MALE: 

-91- 

G.nB PLr.t 3ry?&r R{dell 

Rfid.ll. carad. Enr.,9(12)r 1597. 

Typ.. Gryll$ pbbejq S^wurc 

sFcincN of a lingle ncw sp€ci6 geady lalty wirh thc original 

(R.dell, 1964) of rhii scn6. 

l, n b.Iogrt $ tuittgr.latit q. nor, 

(Fiss. 244-E) 

Sia dtediun (13.5 - 15.0 mm). Body colounlion bl&*ish.bo*n, wirt bNnkh, 

Shining bl&k, wnnour any qmcnrtion: fe bl..k. wirh sligh.eddbh dnge: 

clypeo-fronu I sulure 3rronS ly . ngulat , Oce|lilined with brow.ieh-yettow. Patpibrown. 

maxilldy wirh 4|h sgmcnr shorcr than 3rd aM 5$, th. taod tong6r, widoEd. 

obl'quely trun€t d ar ap.x. Arlenn& blwn. 

Porctuo black, Mowins anEriorly (Fis. 244). 0.E4 - 0.89 rios s sid.6 

pdtfior hargi!, anrerior margnr smnBty @nave. narmwly terrugimus. posterior 

n@ly suaiSht; disk aldo$ glabrols or v€ry fiEty pubenri laE6t lots bl&k, wnh 

iDf.tior nrgin stongly a@nding bekwdds, 

L4s blekbh-brovn, wi6 slight rcddiih ting., uni.olourous, wirho .ny 

odtings; rnleric ribi.. FrfoErcd o, both sid6, wirh a tdgc, Mt, .rcrEl ard a 

small. rolnded inter@l whirhh rynpanuni pdt rioi femon short, strong, tincly 

pub*enq pdterior tibilc dm.d wirh 5 ifi.r.at, 5 - 6 ert rnal poirred 3pici pGt rior 

bai|'Ei wid 4-6 inl.rnal,6- ?€xbrn d€ntict$, 

Elyth slighdy exrending b.yond rh. abdomeo, blow.bh,yelto*, wnh . pale 

latcral band, thc lstar.le cxcrdinS inweil ar te; nnmr diJrirctty wjder do tong, 

wirl idunded olllior nargiri, divid€d behind middle by a curved veh (Fig. 24B)r 

diagoml vcin longi lst chord unifornly fand.d, zod and 3rd 3hnAty siolaE; h.rp6 

with 3 obliquev.iB. sinlaF, Egllsty-ea@d; aph5tfi.ld short. prcvid.d wnh 4 v.iN,

-92- 

trsver'. leirlct forrnlng alMt Ggular, I@ly rccidgul$ celbi laldal fald *irh 6 - 

? r!8ul.rly{&.d @i!4, Sc baritrg I - a bnnch6. wirgr lolg, cddar. 

Abdom .td eEi stolly bLt. Crnirdir (Figr. 24 CDE) ri6 ?itt XB 

roushly quadrat , iE a dior M8in deeply eld broodly emargin E, sidc! iiNat!, wirh 

v.ry fccbly viliblc lal..d lotd. p6l.rio. Mtir witl . r'D[, lpsrrdqild, bluldy- 

poinEd. median lobe; c.6putn!r.s slighly vitiblc b€tl t! .pbhdlu! in (hr.l r|Pect. 

M.@m.rrs (ml: 2 d 

rr.5n5_0 

2.t) 

|lGrr.so 

2.4 

llo|dr?.: I d, unbtcltd_ 

P.nt'pc| t d. unrdEtted. 

rr.73 

o5t 

s.D, 

o.u

w.a. 

-93' 

Eegtfut t6t OlaB {.u.& bar.dFdd, 

dd.r !p.d; B, dtrn di ct, rd!. dr c, .plF.ftr 

d..d !r.d; D, E, pt lll. odd, v.d.l.Id lrL..l .!r.d.

-94- 

BotI mal6 rere akeD from re studenb coll*tion in the Dep.rtn of 

aglicuhlral Enlomology. Universiry of Agricultu.., Faisalabad, Palirrao, 

Dlsrhguhldng clDdsl 

Thir sp@id h6 a d@ @mblu.c to Platsry't" wh@phaus 

subcqucntly dsrib€d by ChQard (1969), bur diffds fton n in $c touo*ins 

Muillary palpi brown; api€l fEld of .lyt *ith regule .ericllarioq lacd fi.ld 

wirh 6 ' 7 veins; abdom.n whol,y bhcti cpiphallus without t unotcd po6t rior 

malgi" (l:ir.24C). 

h is !1rc fairly dislimr frcn P, 8d,eal,r, qkh which its geniblia tu a littlc 

Thc Mm of thb sp@i6 hs bEn deiv€d tron thc criculario. of lh. alical iicld 

of clytn, which is .cgul&- 

Th. rne nat€rial ha bccn d.?osicd in d. la4t Moeum, D.9.rtdctrl or 

Agliclltural Entonology. UniE6ity of Agrhllorc, Faislabad, P,kislan. 

C.nus Crr{rd.! Srusrc 

t874. OryUodes Saussure, Miis. sci. Mexique Amei. enr. Rsh. Z&1..6: 409. 

lW6. (;ryllol4tutrcvu Bolivd. Voy, ltodMhild, lns4tes, p. 196. 

T]4t: edUB ttS LtB \ dk r 

Th. @tlerd mr4rial of a 3ingl. sp€.i6 entircly agrd wiih $c publish.d 

dsriptioi (Chop.rd. 1959: Ou. and Al.x.td.r, l9E3; Sa..d. 1989) of thi. scnu!. 

I edlbd$ ertlrat (tu|}e.l 

(Fig3. 25A_C) 

1859. Acheta t\pplic^ walker, Arn. Mag, Mt,liisi,. (3)4:221. 

t869. CrylLts nun6 Walker. Cat. Dcanapr. Saliit. B . Mos-, l:?14.

-95- 

1869, eryIB p6ttlip6 Wdk r, ldcm, lbid,. lr 51. 

1469. Gryl!.t tiEill4t6 Walker,ldcm. tbid,, l:46. 

t869. Cry 6 suppUcM wallEr,Idcn.Ibid., l:36. 

t814. Grylloda poryi sassurc, Mbs. Mexiquc. ol$., p.420. 

1817. CophoerynB ndlt?ri Sau$uE, M.n, Sc, Cercvq25:233. 

t811. Orlloda p6,!lip4 Sausurc, ldcn. lbid., A: 210. 

t811. erlbds r:gilit6 SaNUrq ldcm. Ibid..25: 210. 

1811- Pi.stodacttl$ ,tlM SaBurc, ld.m. lbid.,25: 531. 

1895- kop.ip.d6 lMinotuB Boliv&, Aii. Sa. .ni. Fr., p. 3E6. 

t899. HoMlosryll$ irr'i.G Bolihr, Ibid.,68: 800. 

t90t. MioSrytLa ttue, ala scudder, Psyche,9r?57 

tq05, GF,llod.s ,@xr Krby, Synof. Ca. Onh..2: /J. 

1912. OryUod$ &r4pr.r6 Chopdd, An.. soc. ent. Fr.,8l:4O1. 

1922. CryIoldtdt M a4$d't r Bolivar. Voy, Rothschild, Itu.. p. 196. 

t925- Gryllod.s aE4i Chopard, Ann. Mag. Mt. Hist., (9)15:511. 

1925- GryAoda rrppr'@ Chopdd, ld.n. lbid., (9)15:510. 

1926. Achao to*to.it Okaati, Dobu|su-Ky@i Korcgi, p.206. 

Thb widcly distribut d spei.s gr.rdy |!lli4 with the publish€d d.scription 

(walkc!, lE69i Chopard, 1969; Sa..d, 1989) ofthi! spei6 exept th. p6t rior tibi&, 

which ar€ wilh 5 spines on @h ndgin (E.ly with 4 or 6on intemalFdgin), nnror 

with 2 dividint veins (v€ry @ly undivid.d) and wings very shon. Edued to IaleEI 

pads in micopr.rous f€nal€s. Gcni6li. (Figs,25ABC), 

M.sultmntl (m): sd 59 

B.L.

,9G 

hlanabad I 9, 8'VIl-C3r 6 d. lljx-94r Pujgli 2 9. &lx_93r Sibi4 9, 

?-lX-93: Zhob I d 3 9, CIX-yJ: Dd llmil Klr.n I d 4 9, 2'lx_9l: Kohx I I, 

4-v!ll.93i Mingoa (sMo I d 4 9, 5-vll-9]: P6haffi 2 d. l-vlll-g3:3 d 3 9, 

2'Ix-9lr I 6, l?-x-94; Ch-gt Manga (Kltuo 2 d, l5-vl-93:3 d | 9, 22lX'94; 

F isalabad t4 d 12 9, lcrv-88; Gutftl. (Feiehbad) I d, 2l-vl_96; H.!d M.tda 

(Sialkor)? I, l?'vl 93i Lil Sohaora (Bduwalpur) 1 d, 2l_VIII_93r Mian}tli I d 

I 9, 4-x-88t Mu.@e I 9, l3-vlll-93i Pakpatar 3 d l0 9, l5-vl_EEi 3 d 2 9, 

l6-vt-88r 3 d I 9, ll-x-88; 1 d,25-v-89i I 9, ?-vll-89; Rlwalpindi L 9,20-vlll88r 

I C,2-lX-88:19, +lx-88i I 9, 8-lx-88i I d I I, 9lx-88r 3 d I I, 9_VIl_93r 

Sl8odh! I ?,25-V-8Ei Sialkol l 9, !5-Vl-93; Badin I d 2 9,24_X-93; Hydcrabad 2 d 

3 9. 25-x-93: sukkur 4 d I 9. 2GX-93: l0 d 25 9, uilabell€d, ntc! fton sMcnG' 

@lletion, D.pdtDeit of Agricultudl Entomology. Uni!6ity of Agdcultu€, 

h h qidely dislribut d thtolghoul PrldsLn atd is ee*ially found in ho$€s 

4s@iarion wnn numn hrbitation, oely @uring in $e leaf-lilt r, *ith 

macropcrou fotns @ll@ld fron lighl!,

-9- 

r,r& 25, Wd.t tuptdN. A1, d t dt 

^, dlob|||a dd.l 

!o.dt I, C, lllll .rara, td.l d 5t r 196.

-9& 

Cryllda eppliM eM e. liSi tt6 wrc prcviou3ly coBidred lo b. No 

diftirenr species by Chop&d (l%9) ($is fauia qs win much bclbre 196?, but 

published i. 1969)- ln 196?, nc aymnyd5.d rhe formr wirh rtr hr4r, bur @ rdyb 

n, Kevd (1960) claiD.d thai O. supplica$ haA prioriry ov€r O. rlsi{or6. 

Ouc (198?b) .6rorcd ihen sht!3 s scpsac speics du€ ro $eir hotorype fcnalB 

deeribed from difcr.rt @Dr.iB and lbe stignr diffcrc{e in hate gcnii.tia s poinr.d 

Thc .bov€ vi.w I de nor hetd good by som. erkd, who ctained rhat d.y 

*er. two lolms of lhe saEe sp€i€s. For .xample, rhc ndopt roB individuh *cG 

produc€d irom $e nicroprercu! parcotl under onro cd laborabry co niom (Chouri 

and McFarlanc, 195?c, 195&, b; TonE, I93) and de by a @mbiMtiotr of f&roB, 

3uch s crowding aod hiSh protlin di€t, rmpenrure, rclative humidny aM day !o nighr 

arcs (Mekiand Otan., 1963i McFstae, t964t Mathad and McFdlare. 196€). 

The larter idea h also supported by the presenr reFarch, whicb r.v€h rhar te 

mrle genitalia in bod lhe forB w.rc exacrl, sinild, 

Kevan (1980) funher slarcd that rh. m&ropt rous rbrms have fu y d.r.lop€

-99. 

1934. Gryltus tanM arsator Uv.a, 86, lo:al. 

19{6. Olltt i[.nt di6 Ctq.rd, AM. Soc. o . Fr.. 109: 164. 

l9!o. Tindbmlla ,4,t ro TadiNkii, Sat. onh. Ic. AErb., p. I14. 

1943. GttIus tnt tut4ta Chopdd, Onh. 4ft. N., p. lE4. 

195t. Grylbt tt tM Rmnq Min. el. Mu. Be'lin, P. 382, 

t954. Acha. @t@ Chc?!.d, v.lh. otlrf. G6. B.!.1, 55: 47. 

$5a. Achrt nnM Krnbag, Olth. Frun. TwkBiaD. p. EE. 

A ein8l. colldt d pan @nforB lo th. publith.d de'ption (Clorcldtov, l97E) 

of rhb so6ck! crc@r tlE ccrotcra@r6, *[h[ e mndly poililil .t th.ir lpk!3. 

Gc.ihtia (Fi8!. 2?ABc). 

Mesuremml! (w)r ld l9 

1.75 

o.r?3 

I 9, qVII-EE, 

;, 

1.2

fit r. lot tfu ,res l'n' d; !, ha d tr-- d, 

|.tal ..p..1 C, .lrln dt D, O. a It I, dlt .b. rlq!

.l0l- 

r\.27. I@ubttjL. bata.. A4, d.r. d, re drhb, 

ddrd q.dr D' C' F[|. ..4Lr, r.dtrl .od lC.l .{d.

aorh the sp.cimeB of thh spei6 werc @lleEd on lighl! 

This spdier has b@n 

_ta- 

rccorded for fie nBr rin. tom Pakisran, 

G.nur rororr.rrlr S.NUrr 

187?. ldo6le,n,,&t Sa!$E, Mem. Soc, Phrs- Hirt. nal. cercvc,25:249. 

lrT€: Luobkw .q@rii SaNur. 

Thc collet d madi6l Fprenting I I spdi.s .x&dy i.llks wnh rh. publisn d 

d*tiption (Chopad, I%9t Saeld, l9E9) of lhis genur, bul diffe$ fron olte a.d 

Alender (l9el) in pr@nting tympatum on both.id6 of dllrior tibir.. 

l. 

(Fi8. 

K.y 

to tle spEl6 ol lrrorLM SaN!rc 

e8m.nt in Mlc will'out i proc€ss or dcmd 

32E). 2 

sgn n! in male *itn a pl!)6 on cxl..Ml aigl. (Fig. 388). 

2. Epiph.llu wirh pcl.rid mediar !06. d@ly norchcd ar rp.x (Fis. 29C); tu|. 

.lylra 3.66lihcs or nor. d long a pronoom, 

Epiphallus qith pdreiior mediao lobe witlDut a dbtirct no0ch at arcxi 

dyh 2.78 rind or ls E lotrB a p.omm. 

l. sia ldge (16.0 nn or more); frcoral rdrum in mle srongly p'ojccrins {nen 

h.rd lcngtl3.z4 mm) (Fis.2EA). ........... -.... r@ai S.|src 

-. Siz! small {15.0 nm or lesti fronhl r6Fum i male slishtly prcJering (nean 

herd l.ngtl' 2.38 nn) (Fig.29A). .,.. -........ .rcolrptB sD. aoy 

Epiphalhs with pqterior nedid lobe alnct ti!&ale or wnh jut a suspicion 

m.dian mr.h (Fis.3rc). 

EpjphallB whh po$erio 

n€did lob€ neith.r ttuMre nor m.diauv mrched, 

1 

9 

l 

5

-103- 

5. Hc5d nlch wider {4.6 nn or norc)t frontd Gt uE mo@ ttd 4.55 dm! rhe 

width of lst aniermlscgn ni .,,,,,,... -...........,.,,.. 6 

Hc.d drcwer {a.3 m o.ls)i frcnt l Bttum 16 tnd 3.65 im6 thc aidd 

of bl orenril segne , , . . . , , , . . , . . , .- .. robrdr.rri, sp, nor. 

6. Hcad *idth 4.65 - 4.95 mm (mon a.E0 nn); clyfr 2.6 tiB a lorg d 

prrtlnwines nuct lo.gq th.r .lylia, , . . . . . . dolidtop.4Dis s0,nd. 

_. H.ad width 5.0' 6.25 nn (Mn 5.71 nd); elylra 2.0 tinci a! long ar 

plonotum; wings nuch shortq rhan .lytr . ......n.lbc.ph4Lt.9,ôr, 

1. 

E. 

Epiphallus wid nediri lobe !ho( rounded at apcx (Fis. 34B). . . . 

Epiphauus wilh median lobe long, fiianguld ar apex (Fig. 338). 

Siu larSc (12 14 nn): lronbl r6rum 

arr.nMl cgmen! hce lebty .xehcd. 

Sia 3roll (9.0 nn); ftodal tGtrum l€s 

.8 

|ffi th2r 4.0 tim6 a wid. a lst 

.......... @r&!6 rP.nov. 

rhan 3.5 rim a. ud. .s lst antcnEl 

rgmcnt: f@ almrt nitt md. 

" l t:":'1':T::TllTll ltl'1 1'll ll lll] lj,H;; 

AntlnnEl plt)6 disdrcdy cuiv.d inqard!. longer $an l5l ..Ennd 

(Fig. l7A). l0 

t0. Anicnnal ptrB ! lidl. longcr thd lsr and 2nd e€nnrl s.Shen$ @g.th.r 

(Fig- 3?A): sennalia wi$ Ndian lobc alno6t ru@le d vcry f..bly chdsiDtc 

at apci (Fig. 3?C)- hanirsqulut sg. no1 

Anre.nal pr()6 much lon8.r ttan l$ ud 2nd antetml egmcot3 togcthct 

(Fig. 388)i senialia {ith median lob. disdrctly nobh€d at apcx (Fie. 38E),

-104- 

r. Ltur.t BM ta.rJ SrBm 

(Fig! 26A-E) 

ttn. L

IE. 2'. 

I@bLtt4, h@rt ]|, baa..dFttu r, rH.p.c 

& dth 6r c+, Fr.n. d; c,.dpr.[!, &,-r grd; 

D, E, DI!||l. &0b, '.dnt ud tar.t.n d&

1,02-r.l l 

-loc 

,,rro.60 

o.r" .* 

ch.hga Musa (K6ur) I d, I5-Vl93t ,auhdrbld (Khuhab) 2 d I 9, l+X-grri 

8 d 9 9, l?-X-94r Mi$! T.9m (Khulhab) 2 d 6 9, l4X-94: l t d ? 9, IGX-94, 

All dE spei,n B of this specilr reE @ll4€d frcm dE lerflitrer und.r fdst 

planiatiois, chiefly of lufaida (E catlrB @nalduL6it). 

A Abdulhn (1955) and S&.d (1989) did mr cpod rhi! +oci4 n ts bcn 

@@rded for the Ii^r time fton Pakiran, 

Msle: 

2, IttubLnût .ucalrptut q, ao\. 

(Fi8i. 294-E) 

. 

(14.0 - 15.3 mn), Body @louradon brown. 

0,95 - !,04 tin s as br@d d prcnorum, fiftly pub*enrr @ip!r 

atd a brud bard on *[ snh, wnici.xleids .nr.riorly !p ro ttE 

@nrex, sligndy ptojeri.e. wirh a traawne y.llo* band bcrwan

-107- 

the lalldl @lli; chck b@n, with a y.llow liNi fu narcEd, rely a Ind. co@ve. 

ddk brown up lo the pctclyp.us, rhc r4i ,lllowishi clypcofbn|.l su$rc st.righr, 

AnErior oclllus ycllow, a.er &o!nd itfably depesd, Palpiyellowbh, the nuillary 

nuch .lon8{., ldt sgn nt longsl, ddLcn d ar aFx .Jd llonB inM Mair. Ani.n@ 

brown, wirh lstsgn.nt rm@(h, wnbour.ny prlEa. 

Prcrctun l].dv.rs. with aloost pdauel sidcs, iE a erid md8in e litde 

@@vq pct rior alnGl rt .idt, both pdidcd with ..ddbh-b@wn brisdBr dbk aln6r 

flat, yelbwnh, mouled eith bNn spots .nd dots, proeid.d eith si&y pub$c€re ad 

small rcddhh brhdes: laEnltield brovo, with a ldge y.llowish spor at ant rior angle, 

exrem. infcrior mrgin bown, iebly endjng pct riorly, 

L4s rrll@ash, mnlcd eith browtr, sttongl' pubd..nti dlerior tibia. pmvid€d 

with a ldgc, crEnrl ald a rmall irt rml rympanunr p6r..ior femoo eith oblh@ 

brown sEize on extcrnal facesi posr.rior tibi& an€d wilh 5 y€llowish spin.s on @h 

side, thcir bss bl&k, api6 brcwd, dediointenEl api6lspurs a lntb bnger lnd$e 

suprior ona. 0.?4 1in* a long d pGcrior bsitarsus. tlE latd provid.! wi|l| 4-5 

inte.nal, 6 cx|emldenticlcs, l6t tanal scsmcnt brcwn. 

Elyra re&hin8 ro rhe apex of abdomcn, brcwn, t.aNlu@n1, wirh prominenr 

venarionr ntrcr loren8.{h.p.d, uodivid.d (Fig. 298); cho.ds lohg, 2nd srongly 

sinuate, lst f4hly roud.d. uniEd with dc Dirc. by l-2 soallveinleE; dilgoml vcin 

almo$ sraigh( harps with 2-3 obliquc wiBi apical ficld ldge, pr*ntinS 3 veiN, 

reticulaiion formed or clongare, alnost r.8uld @lls; IateEl field pal. whiftn, its 

sup€rior n.rgin bown, poviden sith 5 disBndy-phen bownbhveiB, Sc bariog l-2 

brarchd (@cly 3). Wings long, 1.85 - 2.05 rin6 d lon8 a cltra. 

Sl&kish-brown aboe., yellowish benath. C€rci loog,0.8? tim.s a long a 

posterior fcmora, yello*bh. pubenl, with stull bl&k md large whitlh hair. 

Subsenilal plate b'Nnr gcniElia (Fi8r. 29CDE) with p6l! iq ndsin of cpiphallu 

rilobac. ancrior edargimtion broaily V{hap.d. wid g.ndy bisinuo$ m&gis. 

pcterior v*y sballow, broadly U-shaped, bearing a very small nedian lob., the latte. 

deply notched at apexi latedl lob6 acut€ly poiftd and upqardly curled at apex.

Itt. 29. 

LtoiLr'[ .t,&ttut q-e. A, bddFldrd, 

dorsr a.c, 3, drb. d' c-D, 3drd. 3! q c!n.I!, "ord 

.4r.dt D, E, tndr. o!rd, *dEr &d lerl !p.di

I'emale: 

{0} 

It h sinild ro bale, blldifrca in iie folloeing chraccis: 

Frce fla1ien d, a lid. convei- Prorc.un wnD pcteriq M.gi! fcbly @rvd at 

middler dbk srongly 3pou.d wirh brown. Elyrr{ brown, prele,ltlng 9 rcgululy+pec.d, 

oblique v.iB. fably onvcrgi.e rryetd! aper, i.svcM v.illcl! tM@a, foming 

.lon8.t , n@ly @8uhr @lL. Wings 1,94 - 2,06 riB d lona a clyri. bl! tilh on 

sidcs in rcp*, Ovipdilor long, straiglt, with.piol valve nlmw, !n@lh, &!r. n 

Me@.otrdtr(ih):5d 59 

6.201.2 639 

s.D. 

0.53 

16.JGr9.0 

3.?3 0.13 

"." "l

,llG 

ItoldtF: I d, | 7.x-91. lauhebad {Xhuihab,. 

Allotrae: I I, l7-X-94. Jauha.abad (Khushlbl, 

Prratlps: Jauhmbsd 2 9 , l4-X-94; I d. l7-X-94i Mhha Tawana (Khusbab) 

I d. | 9, l+x-c4,2 d, t6x.94. 

Ir ha ben coueied fbn lhe laf-lin . un

,|lI, 

,pically; che.ks b@wni i@ longer than wide, feebly deF€sscd. dark rcddisb-bro*n ut 

b potclt!.s; labrum, mE lyp.u ad tid6 of porlypcu! trllo*irh; clyp@fronral 

suore sraight. Eycs marked with black and brcwnish. Labial palpi y€llow, tn. mdiuary 

brow|li3h, witl 5th scgmnr nuch enldgcd, widening and obliquely rruMl! !t .p€r. 

Anrenna€ brown, wirh lsr Fgmem ptaio, 

Prcmoo febl, {ideniry dl€riorly, with frcir n&gin a litrl. co@ve, p6Erior 

straight. bolh provided wirh smallbrhrlsr dhksligh yoilex, bl&kish-broun, spotted 

with yellowbh, thickly puhcsnri lal.al lob6 bl&kish brc*n, *irh r paL tpor tr 

anlerior anglc. ext cm. inferior hargin srcngly as@nding posreidly. 

tagr ycllwish, lEorSly nottlcd wi$ bro*n; anEdq ribi& p.donled with an 

oval, extemal tynpanun, which is reduc.d to a snaller oft on inlerMl f@i pdterior 

lemora ilickly sriacd eirh bbwo on eltlrnal surfls, darkercd on rpi6r pslerior 

tibi& arned {ith 5 pubc!@m rpiB on @h side, hc lan r brown dd h@l.d at apies, 

black at bs, sup..oinE ml 'licd spuB alndt.qu.l 10 dE niddlc, the l.ror 0.?5 

tim.s ar lon8 a5 posr..ior b6irara; rhe laft.r dncd above wirl 4 i €rnal, 5{ .xrcmat 

Elyr. lhoncr rhan abdon.n. mund.d apically, b@n, *irh a I i8h. hunchl b6d; 

niro! angula. anteriorly, roundcd pdteriorly, dividdd behind niddlc by rwo cuNed 

veiB (Fi8. 308): cho.d! fcbly !i'ule: ha.pr p@nriig I v.iB, $e tBr two sinur., 

thtrd almost stEighit apical field shon, piovided wnb 3 vei6, fotming Ethcr tdge 

tEguld @lkr lateral f'eld ddk brewtr pGenting 4-5 Egulely spa.cd leins, Sc plair 

or having on. bBnch, wings caudale. 

Arown above, y.llowbh bc@lh. Cdci lon8, dcNely pub*anr, bl&kish-brown, 

0.98 rin6 & rong a pclerior f.nora. ccnialia almct sinild 6 L. .AichopMit 

F€male: 

h is similar !o hale qept the follwing.haoftn:

nr !4, 

Itu'.r''& 

d,d*!,{nrr, 

-lt2- 

I 

+-.4Hdl--

-l l3- 

Hc.d ndly 6 wi& tt ptuorunr chelc tlllowilhi ftonr.l crun 3hdr mt 

produced b.twen the ant n@: fae almodt flanecn. Eyd bl@k. P.lpi brcwn 

Prcnoum with sid$ pddlcl, ili pdtorior mdgin fe€bly .iouate. Etvra nuch shortr 

tlEn aHoer, ddk bro*n, witt lidcs lid€r, P@n!n8 

3 fi@ vei6, C! 2'bdEIEd: 

s€ plain, Wings w.y long. Ovipcitor long, alm6t sFaight, with api@l Elvs small, 

feebly flat ned, sn@ti, eul€ at aPex. 

M6!@. s(M):5d 39 

s,D. 

,.;.* 

..;,

25-Vl -88, 

-l14- 

HoloatDc. I d,z-lx-EE, RaMlpindi. 

Allotyp€: I 9, lllx-94, klam.bad. 

Prmtyp6. klahabrd, 29, ll"lx-94; RaMlpindi, I d, 2Gvlll-68i I d, 

AU $e speircB of this rp@iB hiv. hen olleLd fton g'is undd a lighl 

Dletlngulshing cirrocters: 

lr n very simil& lo ,, .lolictoryaait sp. tuv., l@6 snicn n ody diff.6 in lhe 

ddk scrcEl oldntion ard l*r width of hed and fton|'l Bttum. 

Saeed (1989) idcitifred 4 malcs frcn R.walpi.di a L detect6. Btt tt9 

prBcrce of 6 lird on @ip!1, shon brown .lytra, 3 hdp vciB, shon api@l fl.ld a.d 

Sc plain or turdL 3hw.d lhal thcy qcrc nirid.ntified atd &rually bdong.d to ! rcw 

fte me of tis sp€cis hs be.n d.riv.d ftom Polohu. a faoou g€lglehial 

region or Pakis(an. All fie cou@rio. lo.rlitid fdl undei thk re8ion. 

Thc typc rut risl h6 ben dcp6n d in |h. I'gr Mueun, D@.nn nr of 

A8ricol$ral E o,nology, Unive6iry of Agricultu€, Faislabad, Pakisl4 

MALE: 

4, larobl2nnw dourhope^n t sD. nor. 

(Fi8r.3lA-€) 

Sie Eediun (14.5 - 15.5 nn), C.ncBl c.touraion tufaLsladus or y.llowbh_ 

widd rha. pronorun, 1.09 ' 1.24 iM a *id. a pom$n, ..ddilh-brewnr

,lt5- 

@iput *irh 6 short. longnudiralpatc nrip.s, rh. outer rMon 4h sid. joined by $ 

G"liquc lift: doBun wirh a dbtilcr palc @k 0rin8 b€t*@n tc taleEt oce i clGc 

ro $c fron|rl r6t um, $e latler very wid., nildly a.d egulsly @Nq, rilh egutalc 

sid€s: fa@ r litUc @ruvc, l-16 rimes s long as widc, feial rhield browl natdibla. 

labrum. mc.lyFus, sida ofpcrlyp€ur y€llowisli clyp+fre alsun]rc atnoi snaighr 

(Fig, 3 | A), Ey* b@n- Fllpi vcry long, pal., 4th e8mcnr of de nuillary shdt r rhln 

3rd and 5fi, |he lal{.l elorgal.. sronsly pi&nins, obliquely runqr., d&k ned.r aFr. 

Ant€nnae light blwn wnh firsr 2 joi.a d.rk bowi. 

Pronorum slro.gly widetrirB ib front, aoterior rwgin feebly convex, l l? times 

as wide d posrcrior, the latler staighti dhk rranslcse. nonled virh pale yelloq and 

brown, pubese.ti laEhl lob6 ddk brown, with . pale spol at dllrior anglei .xr.hc 

inf.rb mdgin bl&k, a@nding pqtlriorly, 

L-.ss pale. nonled sid brcwn. dlMly pub.cir dErior ribhe pcrfmt d wiih 

.n ovrl, .xrerMl ryhparun, which ir r.dlcd io . sh.ll.r one on inr.rnal f@; p6r.rior 

lbrnoE shdl, rhick, sr.iared eirh bro*n, iil.ruprd wirh ryo longnudiMt yeuos tincs 

on .xlcrnal fae. thei. alic€s ddlq poct rior ribi& arftd wirh 5 dendq ad fcbly 

hook d spiB on @h sidc, n€dioi .rral .pical spus sligndy bng.r rha. 

superoinr€rulrj p6terio. baillrsi fcbly compr.ss.d, provided wiln 3-4 intcrMl, 5{ 

Elytra light brown, witi a ycllowtuh hurcral ba.d, shorter than abdoEcnr niror 

disposed r@ lhe apex, nady 6 wide a! loog, aogulale i. fton!, rou.ded post riorlyi 

divided b€hind niddle by t*o v€ins (Fig, 3lB), vry 6ely wi$ onq chords tably 

roundedr lwps qith 2 3 obliqlc vciNr apiql field shon, p.ovidcd wft 2,3 v.ins, 

rcriculaion formed of a few irc8lld cell3i lareBl ll.ld pale. rspmnr, brewn al be 

atrd along slp.rior n&gin, dh.d wnh 5 lc8uldly spaed veiN, S. plain or omba*h.d. 

Wiry) va lon8, wfinish ar he. palc bown |E at@ 

Ddk brewn on doBun, rllovish bcMth. Ccrci dark brcwn, c.niralia (Fig3. 3 |

-llG 

CDE), with AiphdlB hlviDg V-rlE!.d rli.rix chrgimiid, po.L.iar drrgin 

rilobal.. ncdi.r lob. shon n,|lw, fcbly rcEh.d at.Fx, Lcnl lc'b6 lon8, poinicd, 

curvcd ine;6, benl mar8inJ sinuci c.toparm.rls 6hor( v!ry fc.bly viriblc bctwen 

riuario, t*ra rto ot 

TEMALEI 

"piplalu 

in doBal sp€.t. 

It is sinile io ml. .xcept the following charad.si 

Head rcrf,d, d sidc a prorD$nr fionEl ct ur mr 3troDgl, projerd 

b€lwe.n the ant n@, alnolt ounded; fa@ almt fla! vcry f..bly @nrcr. Prcmom 

pdallcl sidcd, .arcly fc€bly cnldged in tron! ia amcrior rnd po$sior rargi raaight, 

Elytra crF ing !o $. niddlc ofabdom.nr dqsl ficld pte iig I fie EiB, Cu 3- 

branchcd; latc6l field with 5-5 veia, Sc plain. Ovipdilor rFaishr, wit lpiel valvB 

snall, acutely poirt d al apd- 

M.@..s(om):5d 59 

,.ort.u 

,**t 

,,' 

";' 

": ":

e.L.lP.L. 

H.T.L./H.F.r,. 

t.2l 

-ll?- 

t.t+1_23 

Holotrp.: I d, 2GVIII-88, Ra*alpindi, 

Allotrpe I 9, 20.Vlll-88, Rawalpindi. 

P.etyt6r lsldb.d 3 9, L llx-94: P6haw I 9, l-Vlll93r Changa M gr 

(Kisur) I 9, 22-VII| 94; Raealtindi 2 d, 3-Vlll-88: I d I 9, 5-VIll-88: I d I I, 

E-vlU-E8;l 9,9-Vlll-88r I d 19, lO.VIllE8r2I, l5-VlU'88iIdI?. l&Vlll-E8i 

2 9, 3GVlll-EEi I d.7-lX-E8. 

fte spcineN of rhb spaiB wrc @Uel€d from tE l4f-litt r undei dcNc 

plantafions of sufaid! (Eucaltpt$ c@naAubtui!) and other for€st tr6. They wce 

al$ aburdant on lighr h@ culrivaEd ficl&. 

Dlstinguishiog .h.ractesi 

This spei€s lery closcly resembl* L, @ephaL6 and catr be eparatcd 

lrom t by rhc lighl body @lomtion,less width ofhcad and fro alr6t!m, fakly long 

caudat pinsr rnd the epiphallus eith pdtcrio. mcdir! lote @ow, f..!ly Nt h.d at 

The sp@io.a 6lle'.d nigh al.irudcs iion nlow6ire I'rv. ddk r @ld6tion 

$an $os. coll@ted in plaiB. Saed (1989) plac€d enc sp€imens of $is *w spdi6 

nndet L. Mrcb.cphifut due ro $en cbse rcsenblai@ witn thc hroer spaics. Sut 

oi cl@ ad wciul eemimtion, lhey p.oEd io belong to thB Nw !pei6. 

The Mm. of tbis spe.is ha ben dcived from its hiod wings, which are 

Tne type malerial of Ois specia ha b€eo dcpBircd in fi. lnsr M!s!m. 

D.pr(ne of ASricultudl Enbnology, Uriv.Riry of A8ricllrute. Fai8labad, P.kii&n.

Itg. Jl. 

-ll& 

lô .nnB dou.h.f..nk a. w. 

C-8, rEi.!. d, C, dph|l6, rloar 

.&f-, rd!!r rd rrd r.rd.

MAI,E: 

-ll9 

5. latublcMus ne.lio@phalut se. ao!. 

(Fi8r. 324-C) 

Size ncdium (13,2 - 14.8 nn). Ceftdl coloualion b@wnbh,bl&t, r.rely 

Very large, much wid€r than proiotum, 1 ll - 1.25 tin€r d eide s ant rior 

Dargin ofpronorum, ddk reddb[b@wn on doBum, finely plbesnq occipul prcvided 

with 6 shorl t.llow liB, ih. out :r two unn d by m obliq@ lirc on ach side, @hing 

op to the post.rior IEgin of .yB (Fig. 32A); do*um wid a Fav.^. .!ddi5h-y.llN 

band slightly b.low rhe apex of fro al roarrum, th€ lat€r v€ry b.qd. 4.58,5.08 rincs 

a {ide $ l3t anEnnal joinr, febly @nvcx wilh @iDcd .dg6i chek eddish-brown; 

f@ .r@Er.d, | ,20 tiru d loog a widc, d,rt brown. sitn r€ddiih rirgc; hbrus p6lc 

y€llowish: clyp.cftoflal sutuE abse.t !r niddle, ftin8 or sid6, Siviig conov. 

app@aoe {Fig. 32D). Eycs bl@k. Ant rior @Uus whnish, pla.€d a lirue b€low the 

s!.r of ftondl ruhn. Palpi long, y.llowish, thc Millary with 4ih jo'nt dbtimdy 

shoner than 3rd and 5th, th. lltier srongly wid. n8, bownish, obliqely nuMt d.r 

aper. Anlcnna€ wilh firt two eens$ da* brcu, $c Esr lidr brownish-yelloq. 

Promtud storyly wid.ning dt riody, dc@ly pub€$.t, iE .nt rior o{gin 

tuebly convcx, pdtlrior almGt straight, both pNided wid reddih-brosn b.kdcli dilk 

valiegetcd wid brcNn and lclloqishi laEnl lob€s bla.k, with a pal. spot ar infeiior 

maqin @wdds aleritr !n81.. rhi latrr margin endine p6r.ridly. 

Lrgs y.lldhh, rpott d wnh brown, puh6e.ti .nr.rior ribilc prBenring a lrgc, 

ovalerlerMl, asmau rounded interMl tyntanun; portcrior femoo ralhs $ick, skong, 

striatd with b.o*n on erlernal fa6, intcrMl fr@ fe.bly sportd wiih brwn, dlk ncd 

ar api6, p6r.rior tibi& a.n.d sith 5 spiM on bo$ side, @h disin8 iiom a bl&klsh 

pi!, sup.roint hat apical spure al66tequalto medisrpGterior baitai comprcsd, 

da.kcned ar apiB, dh.d with 4 i dEl. 5{ dLbrl ddticlet. 

Elyh slbndy snorur lhm abdoo.o. bl&kish. trarslu@nt, fircly pub.snr, 

chi.i'ly on v€iN; niror M y in apical position, 0,9?-1,09 tim a long d eidc,

-r20- 

rcund.d pdrdidly, oeuh. lnr,iorly, dividcd b.hind nlldlc by l-2 cunrd (rcly 

fu.catc) ycio!; lst chord unir.d qith miru by 2 v€irlcB (ndy 3), oic nd anador. 

the oth.r rM inner ugle, 2!d chod suonSly 3inurcr harps wilh 3-4 obliqu., liNare 

vei6 (Fig- l2D: apid fi.ld rcry shdl, p@nnng 3 v.ia, with. fcw iEguLt ellsi 

laar.l 6.ld povidqd wi$ 5-6 Egul&lyip@d !eis, Sc tuMr. (tucly plair), Wines 

a|&l abovc, yeUowbh-brown beMth, pubscnt. Clrci long, blactbh-b|lM. 

pubes4n, aln6t $ lons as pqteior fcmora. Gcni|llia (Fis. 32o) v.ry crose lo r. 

dolichop@it +, Nr.i epiph^llw *ith dr.rior @gimtion bMdly U-shrp.d. m.dio 

p.sterior 106. a lird. btuc&r, ekally dn6r Eunar., hchl 3idB mor. srongly 

sinuarei ecloDarameB sinild to the lalt r rEci€s. 

Female: 

h t similar to mal€, but difi.R in th. followine ch@l.B: 

Ce.eralcolouration a little lighrcr. Head notat allenlargcd in fioft (Fi8.32C)i 

ircnlal r6E!m shon, br@d, mt prej{ting betwetr rh. .meMr che.k tluowilhi f@ 

flar, yclloebh, febly difiBcd wilh browni clyFGfronnl sutur€ straigh Plonoun a 

little enlarged in front or parallcl sidedi lateral lobB ldSely yellowisb, with a namw 

brown band along 3up.rid Mgin. Elyr..xr.ding to rhe middl. of abdomn; doMl 

field bNi, wnh r yellNisn humeEl band, prdidcd *irh 3 fE reitu, Cu 2,1 

bmnch.dr lateral field prseoting 5 rcellady{p@d vciB, Sc Dlatu or fuete. Abdoned 

browtr dove, y.llowis[ be@th, spo rd will) rhe land @l@r on doBun. Oviposiro. 

long, str.ighr, irs lplll valv6 snall. lan@hG, rcur. !t ar.x. 

Mealreo..ts (m): 5 d 59 

H,L, 

Ma! S,D. 

l].G15.50 14.10 1.03 

r.4t2.25 t.91, 0.22 

3.6S4.45 4or 0.29

-t2l 

9.r4 0.7s 

r.}r.?t 0.29 

0,4J 

6.3G3.r 1.n 0.46 

golotyp.: I d, l?lx-94, Tamb (P*hawd). 

Allotyp.: I 9. l?-lx-94, Tun b (P6hawd), 

Prn.Jp€: lslarrbad I 9, lGlX,94:2 d l7 9. lllx-94i Khuzdar 2 d, 

8JX-93i Sibi2 d 2 9, 7-rx-93r Abbothbad 6 d 3 9, l4-tx-94i 2 9, l5jx-94: 

Maeha 3 d 139, l5lx-94; Min8oh (Swrt I ?,5-vlll-93i P*haw 3 d 3 I, 

2-Vlll-93r Tamab 3 d I19, t?-lx-94; Bhagal (Toba Tet Si.gh) 2 d 5 I, l9-VIIt-94r 

Chanea Manga (kNur) 4 6 6 9,22-ylll-9416 6 lt9,23-Vltt-94r Mith. Takna 

(Khushab) 2 d 2 9, l4-X-94;2d, I GX-94i R.walpiodi I d, ? Vlll,88; I d,25 vitt S8l 

I 9,3{X-EE; I 9,slx-EE; I 9,9lx,88i Ayub PaiL (Ra*dpirdi) 2 d 5 9. l&tx,E8: 

Rawal L2ke (R.walpidi) 2 d 9 9, ll-lx-94. 

rr r very common and widelydistribuEd sp€i€s, col.cted frcm oe tsfliucrof 

allknrds undcrdcnse planhdons, chiefly in for€srs, oichdds, parks and along rcad 3idcs. 

h prefeA hunid pl@s having dense cover of grNes. 

Dini.auisning cn mcte6: 

It hA a cl@ affinity io L. dolichop.rnis sp. m. from which i @. 6ily be 

epa.ll€d by lhc dark colourarion, huch *ider hed a.d fronnl r6r!m, very shon 

winss and se.ilal coifisuratiois (Fig. 32C). App&endy, ir may bc misut€n with 

L. tucrocephalut, but differs riom ir in dE laEral loh€ of pronorun which is with a

II!. 34. 

-t22- 

b 

E 

n 

w 

LrotLlua &.f,ttlg q. -. rl'd.ldFldr 

J, ard rac! B, l!. rrl' lrE l qdr C, rf,. @ 9t 

D. t6 a' & Lr r@l r.d d, r, dth Ji G,.phrdla

-123- 

pale spol ar rhc .nt ,ior .n8lc, .lytra sit! 34 hatp rciN .rn tlc goiialia will! poslerid 

median lobe n*dY lilNat. ar aper. 

Sa@d (1989) id.ntifi.d oi. nale.nd many fcrEt4 ts L. Md.phal6 d@ 

ro th€k close tesemblane qi$ th. laner. B a close dd .dful e&r.iMtion dal.d 

then a a w specis, with th. above difi.enc 

The sp.cimens @ll{Gd al high cltitude have darkct ololdtion $an tnosc 

The mm. of lhis sP@b ha ben dertud fon lhc widd of he.d which i! 

lnterm.diak b.tw4n L. M@.CtulB tnd L doltdlopMit tp Nr' 

The rypc maErjat of thb sp€ics has b@a deposiEd in $e In€t Mueum, 

Departmeri ot Agriclhural Emomology, U riveîty of AgdcultlE, Faieaiabad Paki$an 

6 btobleMut adaut4ws sP. no\. 

(Fiss. 33A-D) 

MALE: 

si& n.diun (l1.0 ll.5 nn). C.reral @lountion blacL. 

Blelq 1.25 1.29 tine! a! wide4 Pro.o$mi eciplt prcsenting 6short, f.eblv 

visibl., yellryirt liBi dcun enh a wrew, |raBrc&, rcddish'v.llry band te'ren 

lalenl o@lli; froffal rosrum very wid!, 4,00 - 4 2 tiE4 the naximun width of Ist 

anl! Mljol.t, f..bly @ns, r.gulely rdnd.d !Fr: chcb su@th r€ddith-brwnr 

face deply cx@uted, da.k brcqn in superiot hdfi tabru6, ant€lvpcu, sida of 

postlt?eu reddlsh-y.llN. Eycs rcddilh-brcw., wilh blick mdkingt Palpi ttllo{ish. 

4rh maillary 3egncnt shoner than 3rd and 5th, tbe latt r nuch widening, obliquclv 

mnotc, brownisn at.p.L A.tcnMe ddk bro*n. 

Promium rra.sv.6e, enldSed in ftont. its adr.ior and Pooterior mtgiB alDost 

smighl, botn prdi.Ld wi.h r.ddish+.@r bristl*; di.k slaeldy onvex, Pub€snt 

blackish-brown, l*bly spon€

-t24- 

Lrgs tellowirb, l&gely spond wirh bNn, de@ly pubcsfl: rlr.rid tibir. 

perfoarcd, wirh a laig! exterMl aid a smll i .mal rympanuni posrcrior remon sniated 

wnh bwn m outr $da6, drkercd ar alicqt, p6lrior .ibia. srn€d wirh 5 inE, 

5-6 outc. rpiM, ncdioiniernal api@t sputs 3liShtly loiga rhan supdiosi 0.8 tinB 6 

long d post€rior b6ilar3B; dic laner shon, deely pubdc.r. ai,ncd abow vith 3 

incrml, 4-5 erErD,l d.nrid6. 

Elytm €xlending !o lne b6s ofer.i, dart bown, pub*e.r in apical p&t and 

o. Eitu; dimr 1.03 - l.2l riM s long s wi&, egltare r ario.ly, reund.d 

posteriorly, dividcd behind middh by a sinSle clNed vcin (Fis. 33A); cnords f@bly and 

reSulely cuoedr h{ps wiu 3 lioure, obliqu. veiBr api@l fi.td sho.t, pr*.lring rF 

v€'ns od a wid. rericuhtior; lal.nl ficld dat bbwr, Fovidcd wirh 4-5 veiB, Sc ptaij 

or fuMte. Wings very shon. 

Bla.l(hh-brown above, yeltowhh benath, Cdci brownilh-yello*, thickly 

pubes@ , 0.9 rins s lorg a p6lcrior fcnoh. Epiphaltu eirh anr.rio. cMgi..lion 

brddly V-shated, with gendy bisinuou mdgin, p6t rior Mgin tritobed, nErti.o lob€ 

short, br@d. abrupdy Miiowi.g lowlrds median apical poiDi, @d apex. e|opeamaB 

witl' apiB b@dly rclnd.d (Fi$. 33BCD). 

FEMALE: 

Ir is sinild io @le ex.?t do following ctratd: 

Hed nornal, 0,99 - 1.05 lidca s wide d promrum; front l rcsrob !ho( 

febly produ@

-t25. 

Ft, J!. l8obbdrut 4gabt8 q, sr. A, drr. ,r &I\ rFrd. 

di E, .rt-&., d6l r.Fi c, D, !a{..ddd, !d.l .!d

1.21 

12G 

s,D. 

"."".* 

"t" ort 

,; .; 

Eoloty!.: I d, IGIX-qt, Ayob Part (R!srtpi!di). 

Allotrp.: I 9, IGIX-94, Ayub pdk (R.walpindt. 

P.dttt€: AFb Plrt I 9, l0lx-94: Blu6a. (Mu@) I d, l3-IX-94; Mittr 

T.q@ (Kh$hab) I d, l+X-94; I d, lCx,94. 

h ha bei colh.r.d Aon.rh. laf-tin ! oadd &@ ptrotb6 of $hj.t 

lEu.atlptas caMAd.,|lO. 

Dldlngnirhlq .brn rds; 

Tn. couedd sp@imc.! k y out ir chop.rd ( 1969) with a. l.rct4d, but dir.r

fton r in lhe follNrn8 chtrectr: 

Frcnul rcslluD not angula in frcnti @iput dmcd with 6 yeuow lifts; chaks 

3n@th: laEml lob6 of porcrlm wnh ! pll. spot at antliior anglq ocrio ribi& 

prB.nring tydp6num on borh silB: cly|ra with 3 harp v.iN. 

The Mm of lhis e@i6 h6 b.an 

cpiphallls, which is Eiangule at rpcr. 

derilcd lroh tn. $reriu n.dis lobc of 

The twe mterial hd bccn dcpGited ii thc Inwr Museun, Dcpanmc.t of 

Agricukural Entonology, Univebny of Agticuliurc, Faisalabad, PalisLn 

7' bto .nnqt .r4t!t!! sP. oov. 

(Fi$ 344_D) 

MAI,E: 

Sir trEdim (12.0 14.0 mfl). G.mral olouanion bhknb'bwn, w,$ 

Lrge, l O? - l l8 rin s .t widc &s dLtio ddgin of Forctun: eiplt 

p.cFnri.g 6 shon loqitudinal y.!lw li6, th. outc. two unit d bv e obliqu. lirc ot 

rhe san€ @lou on ach sidei ftontal rosftm oundlv @ns 4.0 _ 4 2 tin's thc 

nax ihun widd oJ lst aobnnal seghenlr chc.tJ br4!, exftne p6teior h&gin b.hind 

subgcna€ yellowi f@ sliShlly longd than wid., f€ebly e@lded shining r.ddish' 

b.own in supqior pdq @ &ound .ncrior ocellus. labtum, antcclvpeus. sidd or 

pGtclypeus yellowish. Ocelli whitish-y.llow, laE6l two joitrd bv a EanJv€rs' v'lloe 

band slighrly b.hind the ftontal t6rlm. Es broqn, with a fe{ blekish ipors. Prlpi 

ycllowish, thc odillary feblt tinged wnh bown, !p.x or 5rh s8tunt b@n. obliqoclv 

run@rc. Anlen@ b@wn. with lst td 2|d egdents bla.kish-bwn th. formc' 

Prorctun smngly {idening in frcnt, i6 antcrior Mgin f4blv mndve at

-t2a- 

niddh, pmrerior straightt dilk a lnde @nvq, ycllowish, lalgely sponed with rcddbh 

browr. fircly pubecnri laehl lob6 for rh. nsr pan blek, srerir anglc wirh a 

yellow spot, inf.ior ndgin d@idiig pGreriorly, 

t4gr y.Uowish, spod€d wjlh bown, pub6enr; arcior ribia! pcdoErad wirh a 

smau, lounded inner, and a large, oval outer trmpdnuo; pGierior femorn sho( a.d 

tnick. p4nring brcwn obliqu !ri& on dllrEl rid6, brown al r+B; posl! id ribie 

slender, brownish a1 Ocir bsa lnd apic$, &ned wir[ 5 apicdly-h@ked spircs on c&h 

negin, sup..oiiicml apical sp!6 alm6r egu.l lo nedis.0,8 rirc s tong s 

posrerior bdilarsus; thc larer shon, f&bly compBsed, provided widr 3-4 intemal, 4,5 

Elytra a linlc ahoder thai abdom.n. bown, wirh a yellowhh hun€ral bandl 

minor Ery clorc io rFx, aln6r a si& 6 longi dividcd by 2 vciN: hcps wiih 2-3 

obliqu. vei6i apical ficld shott, pEsenti.g 3 v.ins and a few 3mall nr.gular @lls 

(Fig- 34A)i larehl 6eld of rlE se mlour 6 rh. do6ur, pr*nling 5 e$i&ty*pac.d 

veins, Sc furcate. Wings v€ry shon. 

Black abov€. yellowish beftalh, prcvideil wirh a fcw ycllow spo6 otr dorsum. 

Cerci long, d.@ly pubcs , 0.94 ri@ a long s pdllrior fclt@. ccnir.tia 

(FiF. 34BCD) very sinil{ to L. noctuc.phdt8, bur €piphallu qhh ant rior 

cmdgimrion sinuarely V{balcd: elopenerB qirh th.n.pic4 l* poin d. 

M€NurcEors (m)r 5 d 

B.L, 

E,L. 

2.Za

-tzy 

Itoloart : I d, t4.X 94, Mitha Trwlm (KtBhab). 

P.rttr!6 Faqi.iEd (Xold I d, 3 vlll.yj; MrNln I d ' l5lx-gt: Bhtgn 

(Tobr TGI Singh) t d, llvln_g; bl.Mbid 2 d' t ux-gt' 

Ir r!5 f@od in the l@f-littcr undd d.N fordt PlalaiiE ald occrsiodllv otr 

s$$4 undd li8lt.@@. 

DLtltri slini char.deFr 

2.69 

of thi! speid 3lDw clo!. rfiinilv to lh@ of L' MtoctPhalut' 

Th. 8.niLlir 

bur 6 .r.ily b. !.p@l.d by it3 nuch tmattd tit later'l lob's of Prcto|um wih r 

yelloe spot .r dErin .!glc, .ly!: with 2-3 htrF and goiulh with |nrdkr 

cmqrSinrtioo linuilc at|d blcdlv V{h.pcd (Fig. 318) 

Thii rr@icr hs ben Ef.d .ff.r il! clo.. @DbLG !o L' netuc'Plults 

^nd L. rutlloc2Ptat4t +.@Y. 

Th. typc tn.l! ial hs b@n dcpoditcd in thc lnsl M@un' D'Ptnment oi 

Agiiclhuml Entomology, UnircBitv of fudculnlr' Fajehbad Pltishn

.I3G 

n8. J4. L.tabhMlt codu8 lp. or, A, drrn d; FD, |.!ni.rb! 

A' C!6d||l' d.a.l A.dr C, D, ,t llL orrra, rdd rd

MALE: 

-l3l- 

A, Utud.nM duntubt sp. Nr. 

(Fi$. 35A-E) 

Sia snall (9.0 6m). O.renl olouhlion bl&Lisl-bown, wid ycllowih lcgs. 

Daik rcddhh{rown, 1.05 d66 a! wide a prcmruFi @iput wirh 6 shon ycllow 

linesr doBum with a tra.svers band of the same color betwe.n latenl @elli 

(Fis.35A)i fronralr6F!o 3.,14 dm6 d wide s lstail.nMljoint, slishtly prcjdtins. 

wirh mErior mdgin febly 6nvex, a lide rcund.dr ch4k yelbwnh-bmwni faa 

alnosr flart rcd, dut brown in superior pan up ro pGtclypcusr labruD and antccltTrus 

tellowishi clypd-frontal sutlre su?i8hr. Eyes blacL. AnLlior @Uus a litdc below thc 

ar.r of frcnhl mhm. P.lpi yellowish, odillary wi0 a litd. bmwnish ting., 51h 

scgnent cntdgcd, wiihning, obliqNly truMie ar aP.x, Anlen@ bown, with lst 

Promtun trdv.e, sliSldy wid.ning in frcnt. itr .ncrior and pcErior MgiB 

sraighq dist srongly 6nv.x, blekish, wi$ ycllowish spolJ. ycry spaely Pobecntr 

lareral loba blact, *iih a l.qe yeuowish spor along inf.rior ndgin, th. la et sPot 

broadening..Griody, eftar. inferior marSin Mndiry brckwed. 

t!8s y.llNish, f..!ly Aoned with brown; dE ior tibh. p.rfoBcd d dErMl 

f@. with a large, dal rrmpanui, th. intlrn l onc s@ly visiblei Dost rio. f.nora 

elongate, 2.68 tim4 d long a wide,provid.d wibobliqle, bown sfiatiots on eiternal 

tGi pdt rior tibi& dlkcmd al ba aid eci, rnEd with 5 rle.dd, pubes@rr sPiNs 

on @h margin. +i€l spurs long, puba4nt, m€dioi dMb aln6t.qu.l ro suFriq 

ones, 0.8 timer s IonS ar posr€rior bdiralsu, the lattcr. litde @mpBsd, prcvidcd 

{nh 3 intemal, 5 ex€nal dcnticles. 

Elylr. r@[ing ald6t to de .xrr.hity of abdomn, bl&Lish-brown, shiringl 

dimr a lntb bnger tnan widc, angulr in fronr, roudcd pod.riody, divid.d behind rhc 

by a cwed vcin (Fig. 358)i Ist and 3rd chods febly, whil. 2nd sulngly. 

'niddle 

sinuate, l3t united wnh thc miftor by 3 snall veinl€Ei harps wi$ 2 promiient. sinut . 

oblique vei6. 3d small. febly vBible ru_aigh! apiel &.e shon. prcvid.d wirn 2 veiB

'132' 

and nEglld rcticllation; laEral 6cld black, pr*nring 4 rcguldly{pa4d rciB, Sc oF 

branched, Wings shoner 1ha. elytra. 

Black Nbove, y€llowhh b€nath. Ceici blackhh-bown, dcNly pub*!nt, 0.90 

times d long s psi.rior f€nora. Subgeniul plalt lege, bl&h genilalh vcry closc ro 

L. "@M.pha!6 (Figs. 35CDE), cpiphall$ wi$ antcriq e@sination brcadly vshap€d, 

posrqid nedian loh. nuch 3horcr than th. latlBl oH, rourd.d: .clo9&rm.d 

M6uM.rts (m): I d 

B.L. 

E,L, 

A,W. r"R,W 

3.0 041 L55 

1.50 1.15 6.7 

P.L. 

F.R,WA.W' H.WJP.W' P.L./P.W- E.L/P,L, M.L/M.W. H.T.L./X.F.L 

3.4 r 05 0.70 t.tl r.03 0.16 

tlolotyper I d, l0lx-94, Ayub Park (Itawalpidi). 

A snUh mzl€ of this 3p4ies wa @llecrcd fron th€ ldHtue! und.r a dense 

Dlnhgulsl rg .hirrct€6: 

Th. Senilllia of rhis speies 3hN thcn affinity 10 th@ of L. M.Ph.lB, 

blt diffc. fron ir in a nnnber of body che..m, seh a mrch sDilld tiz.. drowd 

b6d ard fronlal dl|m. f€elrlt enldgcd promtln, wnh a pd. spot alor8 infcrid 

marErn of B latsal lobes and fanly bng.lyh, srb I harp v.iN. 

lik.yso in Choped {l%9) b.t*an L. .qucstis utA L. ni8ii..Pr. hdiffcB 

iiom re tbmd in ancn@ with lsr s€gm.nt sm@tlr ad without thc d€ntiforn 

projection and ftom $e latEr in having 6 yellow lims on thedciplt, mnror divided a.d

Ftr. 15. 

,ttotlN4t .'.ara,u |r. E. A iad d ftlrt! d, 

eEr lpdr B, drrd Jr c.Er rEft.r. dr c,.r'sbnd, rtoB.l 

-tdr D, E' t-||] ..dda, dd.r Da L!r.t r.dr-

-13+ 

Tnis sp€cia ha bc.n mm€d aftlr irt v.ry sdrll dy si4, 

TIE holott?e oi thh sp@ier tu b€en d.po6il€d in $e Insal Muelm, DeparrFeor 

of Ag.icultuhl Ent mology, U.ircGny of Agftultur., Falrlabqd, Pakk|!n_ 

MALE: 

9. IirobbMat pilk na sD. aN. 

(Figr- 36,{-D 

Sia sm.ll (9,5 - 10.4 mm). Body @lou6rio. Gddish-bwi, wirh lcgs rt owish. 

Reddbh-brcwnr aciput qirh 6 rhort yellow liB, a rasv.Bc bard of rhc s. 

@lour *t nding b.reen lhe lal.ht @Ui sliSlrly bchid rhc ap.r of fionnl rosFun 

(Fig. 364). th. lartci @r vcry broad, 3.75 - 3.91 rims as wide a tsr anrlnBtjoior. 

almcl ruMre, with rourdcd anglsr chelc brcwn, with yettwbb lin : faa ddk 

bown in superior part; l.brun and $tr.typeur yetlowi.h; ctyp.cfrcnbl 3ulure ,tr-aighti 

pmElypea wirh sidcs f4bly p6j.crcd posrcriortyt a@ b.lwen ancnnal s@k rs and 

bas of nandibla wirh f@bly depr6s.d inpr.ssione on borh sides, Ey6 d.ik. Ant rid 

Gllus y€llowhh, aB around it f.ebly deprc$ed. palpi yeltowish, vith 5rh ndjuary 

$gn m d{k.ne

-135 

long d m.die oB, 0.9 rin4 s tong 6 p6t rior bsirarsu: rtc laftcr 3hon, f€bty 

cohprs.d, provided with sho( deidcle3 and tongef.eddish brcwn bri$tes. 

Elyra paL yellowish, ranstu@.t. .xr.'{iis a tide beyod ih. bsb of @rci; 

nirror slightly longer $.. br@d, undivided, ansular! dl.riqly, i.tlly ,ounded 

p6Eriorlyi lsr chord rlighlty @nded. joining lhc mi@r by l_2 vcintcr!, 2nd srongly 

sinuaE, 3rd almost sFaighti harps wilh Z-3 slighrly sinuate, obtique veiN; api6l fietd 

wcllne!.lop€d, wirh 3 veiN, rcricutation v..y ieal atm6r E8ula. (Fig, 35C): tsr.El 

Iield palc whirish, wirh a Mrow brown band ltong supsio, rhegin, arned wnb 4 

dhh veins, Sc on.-b6rched. Wings very long, t.B-2,2j tim.s ar tong s.tyrB. 

Bown aboe, tellowish b.@$, Cdd tong, y.uowish, rhicktyput sm,0.95 

trD4 a long d posl€rior feFoB. ce ratia lery sinild lo lhor. of L, hmihamut6 

sp. @v. {Fi8s. 36DEF). €piphlltus wirh aoteior emalgiulio. v{hap.d. pcl.rior 

mcdian lobe sho.t, n*ly lruneE or tebty mrclled at ape. 

Mesunmentr (mn): 4 d 

t3.33 

r,3l

I d, GX,88. 

.136- 

0 56{.63 0.60 0.01 

3.53-4 19 1.31 0.27 

l.r?-r.3t t21 0.05 

0.714.16 0.01 

HolotrD.: I d, 2qlX-E8, coja {Faisalabad). 

Prralyps: Faislabad I d. l6-vU-8E; Baharalpurr I d,25,VI-8E| Sa4odha 

All 'lE 3pccim.N of rhk spei6 werc olle.Ed fron lighr in culrilar.d fi.tdr. 

Dlsrlngubhidg chlmler.: 

k hd r cloi. r.scmblare ro t. rnkm.r',6, bur difi.B Aon ir by rhe a.r.ml 

pmcss, which i5 lmalld thd the lcnglh of lst snlen@ljoin! miftor undivided, h&ps 

wnh 2-3 obliquc vcins and m6!En nls of krioc body parll. 

Thc genitllia of his spai6 snov dgn {finlt! to lt@ ol L. h.nihtwlus 

sp. iov., frod which it diffeB in much shoner anrenMlpl!)B snd fronl.l6rum, the 

l.ner not .pidlly thickqEd, wi0! a traBv.rs bdd a linl. bchind i(! ap.x and f..! wnh 

fccbly proj@ting tronbl roshm. 

This spe.ies h6 b@n inconecdy id€nlined a! t. oreuatl$ by Sac.d (t989) due 

io its clos. rsenblrc with lhc lar.r. Bur rhc abovc diff*n6 show rhar n was a 

cl@ly .elated ncw speia. 

This sp@ics ha b... un€d be!!e of i|3 difiere r@s fon r, i .tulirc and 

L. hdihatulu sp. N. 

Th. typc mateial of lhis speci.r hd ben defsiEd in rh€ lngr Muscun, 

D.prtncnr of ASriculrual E ronoloCy. Univ€*ity ol Agricultu.c, Faielabad. P.tbDn.

-13?' 

fl 

fi 

wI n& 36. Loata.ffit pln ru rp. Dr. A, h-d rd Fflra a, 

&ril 4dr & rt .dd *a!d 6t C, dttr. dr 

Dl' *lh r; D, Crbh' .h.-l 4dt E r' ,bl[. 

oqaai rir'l Da rari r{d.

MALE: 

-t38- 

10, htoUehlkus h.athanulus sp, aot, 

(Fir!. 374-E) 

Sir smll (9-25 12.00 nn). Bod, @l@Etion y.llowish, veied *ith brom. 

Slid y wider.han prcno{un, 1.03 - l.2l rin4 the dlerior width or dE lnt r, 

wnh doBun brcwli 6ipd with 4 ycllow liEs and a wid. @t$ bond of the sNm. 

colour, $e l.rter extending antcriorly up ro th. polt.rid nargin ofeyes; fiont l t6F!n 

widc,3,60 - 3.92 rins d wide a lst artenMljoirt, ib apidl ddginslightly thickcmd, 

with a raNvere yellow baod conn@ting tbe latctal Mllir chelc brown, with a 

yellowish line: tu e shioi.g brown, whh labrun .nd ameclyp.us y.llow, feial thjeld 

Uar, wnn sdongly proj4ting frontal rosrun .M f..bly @nvei sid*, @ bctwen 

anl.nEl @k ts and bM of nandiblB wilh . d.cP inprNion on @h sidci clvp.o- 

liontal sutue straight. obtuE at niddl. (rc.y naly d..ply €mbedded), {ith both iida 

proj@ting on p6rlr.6. EyB bl&kilh-brcwn. Palpi y.llowhlt, 4th mdill..v *gd.nt 

dasri.cdy sho.H tha 3.d and 5l|): th. lattcr dTl.Nd, obliquelv ruMEd ar aper' 

Anlen@ *ith Isr Fgftm bwn, cxchal.i81. vid a elaivclv thql lhin bwnish' 

un*gme €d, slctrder pilx*, *hich is inwdlv cuNed tikc a hoo& (Fi8- 374). 

P.o@tun srongly ratsv.*e, slightlybulgiE inlhe diddle, wiih aoErior mdgin 

tlcbly @neve at middle, poste.ior sraiSht, boih nnelv liE! wnh brownr disk a lide 

convex, colered witb silky Pub4cence and long ruious bristlg vcllowish *ith a 

m.diah brown lirc and nunetous b.own rpoBr laErd lobes brown in supeno. Pdl 

whirisb i. infdior pan, thcir irf.rior marSiB bown, al@t shrght. 

t gs yellowirh, nonlcd with brcwn, thicklv plb€snr; ant€rid tibi@ p..rorer'

-139. 

Elylr! long. trdl@n! €rEnding ro dE apex of rnbSpdit l PlaL. y.llowirh, *ilh 

promircnr b.own EiN; nimr sligtuly longd thd widc, aryul.t in frcar, h.!ly 

.ouodcd pct.rioily, undivid€d (Fig. 378)i chords long, lst at

2_0t2.15 2.Zt 

-l,lS 

Holottp.. I d, 8,VIt-9t, htadbad. 

Alloty!.: I 9,8-Vll-93,IstaDbad. 

Praryp6: t3'am.bad 5 d, E-VI9li perhawd I 

(Faielab6d) t d. 2l-Vt,93; co.i6 (Tobs Tek Sineh) 4 d 2 

r.r lx (R: I rhnre I ,l . 25 Vl 88: Mi.nwiti 4 ,1 2 9. 4 X-BB: 

Sareodba I d, 5-X-88: 2 d I 9, 6-X-88, 

l.atr.3t l_7r 0, 14 

2.e|1.0 2.41 0.t5 

5.n 0.53 

r2.614.m 13.33 0.?9 

Thn speie is connonly found on tight !E cultivaEd fi.l& md forsr 

plan$tion!, but nr.ly in lne taf,lirtf under deBe pl.nhtDB. 

Distingoirhiry chaEd€n: 

3,gB.n 1,77 o-to 

1.03-1.21 Ll2 0.06 

0.55461 0,J9 002 

r.35-{,00 1.12 0.21 

r.o 0.05 

0.7r.0.30 0.?3 0.02 

d, 2-VI[-93i curMla 

9,2E-tX-88i2d39, 

rr hs a very clc€ @nbtae |o {,, /@&U.a bur qn b. epdated by i|3 

smaller body sie, ahrertul prdH3 and eeniralja wirh nd,an posLnd lobe atndt

H 

A 

\d 

-14t- 

LbtLw rdit utls 4"i.4, tirlElqld 

d; B' drrn itr C-t, !..rdir a ; C, .!qntr!, dodt rp.d; 

D, E, !6.llL u*x, r&rr .nd Lrd r0..l!.

-142- 

Sa@d (1969) idc iticd onc mal€ rron coja dd 3 mla. @h fon Faisalabad. 

cojn md sdSodha, .s L. eque$ns ^nd Z. dcorr Bp@rircly, bur ! c{re{ul 

ex.miution of their aniennal proc!$ and o6er body chtrscren rcv@lcd that both the 

specia w€re miridentificd duc lo lhe brekc. dt nDl p|6*, .!d acnElly all the 

srecnrens wcre of this re* specia- Ir wa lurrhef slppon€d by th. dbsecrion a.d 

adltsb of tlE balc gcnitalia which *!rc quiE diff.mr iiom the larcr speiar. 

rhe saFe autl'or (l@. cn.) placed dro@ully a lars. numba of lp@imcB ondcr 

L. h@liJ.t, bot tE, tte bcloog€d ro thi! ew spei4 on $e bais of th. abov. 

Tt. nlne of this spais ha beD 

snaller than rhar of !. /larrly'.r. 

d€rivrd fon lh. ancnnal orlxB. which i5 

Tbe type dat rial of this sp.ci.! ha be.i d€pooircd id the lEcr Museun, 

D.part@nt of AgriculluFl E ohology, UniveBityofAgricullurc, Faisalabad, Pati$ar, 

rl. Iitoblehd8 honuwer CboDtd 

(Figs. 38 -F) 

1969. LoroblwN /t@ll.r Chopa.d, FauM of India, Cryuoid, p. , lzE. 

1949. laoblM Jinahise\ M.&, rh6is. Univ- Agric. Fai!.l.bad, Pak.. p.88. 

fi. couel€d m{t ri,l of rhh sp€cies grady ag.6 with th. original d.s$iprion 

of Chop&d (l%9) and subsqu.nr des.riptior of Sae€d (1969), but dificB in thc 

L.8s pal., yellowish; elytra widi l st choid uniting dE nnrcr by o@ s@ll veinl€t 

(@ly wi6 2) (Fig.38c): wings lorg,2.0- 2.2Elire 6longsclyrai gcnir.lia wirh 

ant€rior edargiDlion bradly V{hrpcd, pdtaior dedian lobe dirrimrly mrchcd 

(Fi8s. 38EF). 

TEMALE: 

Il is a rew allott?e qhich r*nbl6 .n! mdc, bur difr.B in rhe following

-t43- 

Had 0.95 - 1,03 line! s wide s pronotuni liontat rostrun shon. febl, 

conwx, uniformly roundcd, mr uEriorly prctong€d ard rhictcncd ar &.r c[.et! 

ycllNishr f@ flatr.n.d, wiri labtuo, ctyPe8, nuldibt6 yclwish. er brown; ctyp4 

frontal suturc a linlc @nvcx in thc niddt.. Ant nMc wnh r3r segnent smoolni witholr 

any dt hrl pc6s. P.orciun eirl po6crid Ddgin fe€bty sinuE. Etytra ercdi.t 

to lhe bes of erci, wi$ 8 resul&ty obtique veiir, firsr 3 fre, [aDsvcrs veinld! 

nunero$, fordilg.tongai. @lk (Fis_ 38D). Winss tonc, 2.08 _ 2.34 dm.s s tong 19 

.lyt a. C.rci ycllowirh, pubesc.nt. Ovip6iro. tons, snaight, with rpt6l vatvc. sm.lt. 

utrow, la@olale, acu0e ar apex, 

Msun@.c(n!):5d 59 

Mrt. 

,.rr--r.r, 

lr.sGl6.50

ni. 3t, 

btotLn$t lwslrq. r, lad.d Fldo 

a, ,b.td 

{o..ar a, ir drdr .td.dr c, d}tn ,r D, tt e l; 

E, C&rr, da.r 4dt I, !bo. oeda, r.E l .4d. 

D

_145- 

AllotyFr I 9, &x-EE, Faiel.bad. 

Pmrypes: Islamabad 2 d l? 9, 8-vtf93i Abbolnbad I d, l5lx-94; 

Bal'awalpur I d,29lv-88r I d,29'v-89t F*alabad 1d l0 9, 1GVII-88: I 9, 

r9-Vl9Ji | 6, l+y194:2 6,2!VI-93; Goja Oob! Tek Singh) 5 d 15 ?, 2&rX-8Ei 

l2d 15 r,29JX-88i Hc.d Marala (Sialko, I 9, L?-VI-93; L:hoE 1 d,25-\.1-E9i 

kyyah 2 d,29-V-89; Mimsali 6 d 6 9, +x-88; MDfhrgeh 4 9,20-vII-881 

Palparbn I 9, IGVI-88; I 9,E-vll 88; I I, qVIl,88: I 9, IGVII-EE; I d, 4-X 881 

19, ll-X-88i 1d 2 9. l2-X-E8; Rawdpindi I 9, l7,VIII,88i 2 d 19,20-VIII-88r I 

d, 3lx-88:ldllq8lx-8E: I I IGIX-88i Sdgodhr I d I !. 5-X-88; 5 9, 5-X-88- 

Hrblrtt: 

It h.s been couectd mainl, on lights @r cukiuted fields and forsl plsladoB, 

bur.ely in thc l@fiioe. undr fo6t plurltioN. 

sae€d (1969) describcd a holorype male from Ravalpindi 6 L. jitushi, bll 

a@lrsis of $c anEnrai proc..rs snd m.lc gcnilalia prevcd i6 Disiddnificarion drc ro dE 

brok.n antenEr pr@ss a.d proved dat ir &tully bclong€d !o L hanalila, tercfot. 

synonymiz.d wnh the lattr, 

New auotype femalc of this spaie hs beln dcsdib.d for rhe lml tin. fron rh. 

typc l@lny. h ha b€en rccordcd lor thc fiBt dne from all lhe abovc t@.lilict. Edlier, 

Chor.td 0969) .Lscdh.d it frcs Faisl.b.d s'd sae.d (l 9E9) @lleI.d fton F.ist b.d, 

Cojn, Miewali, Pakparran, Rrwalpindi and Sargodh.. 

TIE auoryp! feoalc alongwitn odEr speid.N ha ben d.pcied in rhe tngr 

Museum, D€parthcbt of Agricrlrural 8nlomology, UrileBiiy of Agricult!rc, F.isatabad, 

GoN Y.Ldt rod Rfude 

19n4. velsili.totu Randcll, C!Dd. Enr.,96(t2): 1s86. 

rW. S.apsip.da n.ad, SaNuc

.t4c 

The colle.Gd trE&ri.l belonging io 7 steci.! q&dy r.llis wirh rhc original 

dcscriprion of R|ndell ( 1964) and subscquent dscription of vrl{rt .t ol. (t915) ot thits 

scns, bur diffcB fton Ou. rnd Clde (1983b) in lh. anErior tibi&, which ar! pcrfor.r.d 

on both sid6, .lyh i. n.l6 @hi.g alm6t !o lhc ap.r of ahdoo..! h fcmt6 neh 

short€r and in r*urenent! of veious body pdts. 

Key to tt sreid of Vdulfdttut Nn&tt 

l. Had with a lraBvlM yellow linc or band bctwdn la,r..al@lli. . , . 

_. Head wilhoul a ralavm yellow lire or band betwEn l.chl @lli. 

2, 

l. 

Sia larg. (12.6 l?.4 nn)i postclior tibiac lonS (6.0 ' ?.7 nn): Mndiblca long 

(2.75 Dm o. nore), uy prcniEn1. .. -..................... 3 

siz. snrll (ll - 12 nn); pdrcior tibi& shon (5.2 5.5 m); frldibld $on 

(2.25 mm or l€$), wnhout aiy sp*ial @nformation. . . . . . . . . . . . . . . 5 

H@d sith a wid. ydlw bad b.tGn lat 6l clli; h@ .bruprly d.pl.s.d jBt 

b€low dtc aDlerior oceuus (Fig. 40A): .piphallN wilh posi.rior mcdi$ lohe les3 

p@dued (Fic. {OB). . . - -.. - -.. - -..... - -............ -.. 4 

Hdd eith a n.llw, feeUy vtible, reddbh{rown line b.tsE! lar.6l @lli: 

r@ abrupdy dcp.Bed clype,Iioocal sutue (Fig. 394)i eliphallu wior 

pcr.rior neditu lob. di5rimdy prcdued (Fis. 39C). 

Sia ldsc (!3.? - 17.4 nn); @iput d{k colourcd, with 6 yellow linBr 

prcrctlh enh hEral lobes conplcldy rlatk brwn, *ith only a 3tull ycllow spor 

at th€ ant rior an8lc. . . . 4qpdlr (Walkei) 

Si& 3mall(12 - 13.5 nn)i @iput liehl @lourcd, wilh 5 brNn ban&r prono$m 

vnh hErd lob.s drk bNn i. supdior pan, 6t light @louEd ilonS infcior 

margin, ,,.,,.,,.,..., lunlrotut sp, nor.

5. 

6. Miror uodivided; .lrta wnh lat.hl fi.ld p!le, with a br{M 

meSiii AiphaUus *ith porr.rior ncdian mtch 

MALE: 

laqe (Fic. 44). 

-147- 

Hsd wilh a wide yellow b.nd betwn laicBl ellir @ip light @lourcd, viih 

5 bnwn bandsi @iphallB wirh p6r.rior mcdian lobe shon, sliehrly mrchcd c 

apex (Fi8.42C). . . . . t ti4tud6 .9. Mr. 

Hrid with a very '.'jw, feebly visible, yeuow line b€iwc.tr l*.ral @llii 

@ipur dtrk olou'!d, pNidcd with 6 prcEiFn ycllow liB: .piphalls with 

Posterior nedid lab. lone, lljghtly .nrgiME at lpcx (Fie. 4lC). . . , . . . . 

.omi.a,i sp. nov. 

elyre wi$ lNrr.l ficld llDlly dek brcwn; Aipndlu *ith 

noEh and lat Bl lob€! small (Fig.45C). . divtdlt .p. nov. 

r. v.ta'llictotut 4rr.o16 sp. dot. 

(Fi8s. 394-D) 

Sizenedium (14,Eno). Body@l@Etiondakbtown, withle8:liSIGr,sF .d 

Dst blrckbh brown, wider l]lnn Drcnolumi @ipulwnb 6 vcllow lirct' *nh a 

very nurow, iirely visible, median lirc, thc outer 1wo aln61 obliqu. @chi g 

ankriorly !p !o rh. pqterior Mgii of.y.s; fot.h..d proi@tiig belw*n st nna., 

abulptly deprssed nd cl}lcet onul slturc (Fig. l9A); f@ wilh slperior Pr d.rk 

shining bmwn up io p6t lyp.Lit, *ith a ycllov spot at dlddle ju$ abov. the &pGsionr 

chc*! fof ln. d6l pan yellowbh, d.a bclow .nd bcldnd eh eve b.wn. Evd sPo&rd 

wiu bt@k .rd r€ddish-bwn. Oc.lli y.llowilh, dE lat Bb uiEd with a rcrv nlrcw. 

.cildirh-b@n liE, which is fcbly angul.t !t middle Maldrbla! labruB r|d 

{n6ly!.s @mpletely ycllNish, th. fom.r {tongty elongaE (2-75 nn) naking he2d 

2.0? tin4 d hign s Forctun, PalPi y.lloq, 6.sth Miultv brsd.ning obliqu.ly 

trln€tc, with a lighl bownish tiiSc at aP.r' Arl.n@ brcsn

-t48, 

Promtlm FaNGe, mrch widd rhtn lon8, .nlargcd in fiom. irs di.rior firgin 

fe.bly corcave, posldior 6loight, both provid€d with dcM re* ofbrkrt.si dirt3lighrly 

conver, firely pub6&nr. povid.d sith ycllwish and bla.kijh spo6, eirhou ey r!@ 

of Eddish ti.gei laEral lob4 for the most pad yellw, in srp€rior maryin blackish- 

brown. qtd. infqior D&sin blek, sliddy ending ponerioily. 

LrSs tellowisn, sponed wnh bowni anE id ibi& prfonr.d wnh a ldg€, owl 

crterMl. .d a very oinlle, rolnded i.r..Bl .ympanunr pcLrior fcmod elongrc, 

obliqu.ly striat d qilh biown, th. la$er inremprcd enh rwo longitudinal y.Uow tiEs: 

pct rior tibi.. med wilh 5 - ? inr.rnal, ? , 8 dcrDl, y.llowbh, apicdly curycd 

spin*r post€rior bGit si wirt 5 inl€rul. 6 err.@l s@lldenticl.s. 

Elyfr brcwr, rmluent, .lb6t t@hing to lhe bsls of ercir nirw lozcig.shap€d, 

divid.d o@ly al niddl€ (Fig. 398)i chord! slightly cuN.d. unir.d with ni@r 

by . sinSle vcinlct .@ lrl.rior angl.r harp6 wilh 2 wc ndebpcd, sirule, obtiqE 

verc. aod a small, alnosbfaight rein, {hich app.rB ro b. a bEmh of 2nd haIP; lpical 

neld shon (l.l mn), povided wid 4 vciB, relicula,rion forned of long, nar, rcgllar 

@lls: l.tc6l ficld paL, wi$ a ddk suFrior band, previd.d wirh 6 .quinLbfi, @gutd 

vei6. Sc b@ing i , 2 br@h6. Wings !hon. 

Blackilh-brcwn above, ,lllo*ish benarh. Cerci brewn, sr.aighr, clor€ ., 0,9? 

rin$ a long 6 pccrior f€nora. Epiphltls witl| erc jo Ddgir darly aln br€dly 

cmargrnate. lne pcterior trilobatc, *ith ncdh tobe b@d, odpuatirety tong, dc.rly 

,plcn€4 at ap.x: etopemr4 pcEridly forhing long, heklike srlErE. eirh 

internal n&sinr almct rounded (FiBs. 39CD). 

Mslrctlenrs (m): I d 

B.L fi-W. P.L, P.W. E.I-. M.L. 

11.30 4.70 2.50 4.50 7.3 1.95 

E,L,/P.L, 

M.W. H.F.L H.F.W. H.T.I,. 

2.30 9.40 3.15 1.7 

M.LJN'.W. H,T.L./H,F,L

t. t9. 

-l49- 

D D 

V.brli.h@ dr.tlat .0. u. A, Li dd F{.6e d, 

L-tl .+dr I, drr. 6r c, .!*L. &rt edr D, ,a.n&

,l5G 

HoldtF: I d, l5-tx-91, Maeh6. 

A single nalc of this spei.s ha bc.n @UE|.d f.om a dia fEtd having d.nsc 

Dbtiryubbiq .h.n t€Fl 

Thn spei.s !ho* iE d* afii^$y t/t V- Bpqt6, bul diffcB in 0E fa.ial 

deprc$iod, laG6l @lli conmcEd wirh avcry narbw tinc aod th. genilat onflglEtion 

(Figs. 39cD). 

fie namc of lhir speies h.5 b@n deriv.d iion i|. f&i.l d.pKion, which 

abrupdy shrb n6 tbe clyp.efronul sutur€. 

The holotyp. naL h6 b@n dcpositcd in tfflcr Mscun, Depetment of 

Agriclltunl Enromology, UniveBiry of A8ricllrui€, Faislabad, patb{!r. 

2, Veta4lictotit aqp./slr (Wrtker) 

(Figs. 404-D) 

t869. Grylrs 6pqsa Walkcr, Cat. D.mapr. Satat. Srit. Mus., 1r39. 

1E11. Gry[oda ,.rtu46 sau$uE, Mcn. S@. C.mvc,25:205. 

l8n, Scaplip.du urz&lr!&tr SaNue, Mm. S(x, crey.,25:24'. 

1924. GryIodcs 6?, 6 Uvarov, J. Aliai S@, Bergat, t5: 3l?. 

t925. G.tll6 b.nh.lts Choprd, Am, Mag, Mr. Hht., (9)15:510, 

1910, kaptipeda ap.6d Yin Chi Hrr, p€king m. Hnt, Soc, Bult., jr 22, 

1936. GryAulls 6rvn6 Chooard, Nota3 Ent. Chimie, 3:6, 

t964. velatif.totE 6p.dd Rrndeu, caMd. Enr., %(12): 1587_ 

1964. Velorili

-151, 

Prcnotal latcol lob6 brcwn, qirh a pale spol !r.ftrior lower aogtci mir.or 

dividd by a cuRed vcin, Sc bdjnS 2-3 bnnche3i ferulc clyra enedinS !o rhe ap.x 

of 4th abdomiul t lgit : wing! v..y shon in borh sx6. Gcnioli4 (Fi!3.40BCD). 

Me&rurenents (m): 2 d 39 

ild. F@L 

l6-x-94, 

H.blttt: 

3,33 

7.5 

,,rro., 

9.Gr.3 

r,52-2,t 

1,0t.t,22 

Artb Park (RasdpirdD 2 d 2 I, lGlx-94i MilE T.we (Khushlb) I 9, 

All thc sleirctu of dris spe.id *r. coll{cd fton O.l@fiird lrdd dea. 

fb.csr plantatioB of s\fîd^ (E@otpt6 cmldllrBtr) aid si.sbao 

1.20 

This spd! ha ben @ord.d for rhe fiBr dne frcn P.kb!an. 

";, 

; 

",,

I[t. 40. 

-t52- 

c D 

V.lstfi.bntt otaqfut. a, !.d.nd r.@to d, hrdd 

.r.d; l.D, Frd. d;r,a!5t la, ddrl4dr c,D,![.llL 

...d-, r-!.1 rd L-J .,t rlr.

MALE: 

-153, 

l, Velat .to@ tu/iaorb 3p. !oy. 

(Fi8!. 4lA C) 

Si& ncdiln (12.2 - 13.5 nn). C€nehl colouBtion lighr brow., wirh lcgs 

ycllowish. nore or l€ss sport d rilh brown, 

Dark brown, mwh widcr rhd pronotun; occipur lighr @lour.d, p@idcd with 

5 brcwn bands, rhe olrer lwo snaller rhan the iore6; loEh€ad prcjccring betw@n rhe 

anlenm, rbruptly dcpBsd slth{y bclow rhe n.dian @lla (Fig, 4lA), rh. trttcr 

giving bdry a f€dly vjsiblc wow yeuow tinq che*! b@wq virt . paL aq b.hind 

cy6 aod on 3ubg€E€i l@ ddk brown up to rhe middle of clypcu!, liehr brceo on 

an|4lt?cls and labrun. Ey4 bl&t. Oc.lli FllN, ih. larcnt rm uiired with a palc 

t6Nvep badl. MadiblB jallN. sdongly lergd.!.d (t* rh2n V- otp.Mt,2.75 - 

3,25 mm longi palpi palc, the mdillary wir! a IiSIlt bowtrth dngc, 5th s.gncnr 

le.g1hened, widening ard obliqu€ly trundre at apex. A.t rn& brown, wirh lsr s€gftcnr 

pde, spon d with browr. 

Plonotln wider $ad long, 0.56 - 0.62 tines a! long s wid.. ib derior margin 

febly @Mv., pGterior almGr strai8l , borh lircd with rcddish-bio& brisrj6, iid.s 

rcbly bllgan8 out in niddl.; disk monled wi$ brcwr spoa ad dors, pubdenr; 

larcnl lobcs *ith superio. pdt ddt 'rllow, brwn, inferio.lighr colould, Ue qFeoc jnfcrjo. 

Dargin bl&k, slighdy ac€Dding b&kwdds. 

143 y.llwish, dcly spotr.d wirb brcw!; &r.rior ribia providcd with r ldge. 

o$l tympdnlm on exl! nal face, the inr.ro,l havi.g only a dcpr*ioi or rcry lmatl, 

febly vhiblc, .ou.ded oDci posr€fior f.no@ sho , slout, srir€d with brcwn on 

crbrnal f@i poslerior tibi& &oed wirlt 5 lpicauy-h@k d spiB on .!ch |ndgin, eilh 

th.ir bas rnd apie d!l; p6Eior bsillrsj a lnd. @np6ed, provid€d win 5 - 6 

int na!, 6.7.xt rnrl d.trti.lB. 

Elyt! c.tiEly lighr brown or donun; Diftor weu-d.velop.d, rcund€d 

posleriorly, divide

tj' ar. 

-ty- 

E 

Yl|{/r4,s tu+-.AHd:r-a, 

td o.q l! {h dr c' i*l' &d rd

.155- 

the anterid mgL, 2nd .hod sEonsly bcnrr haQs with 2 obliqu vei$; .pi6l tEtd hrtr.r 

shon, pdidcd wir} 4 Eins, forniDg alMl sq@, idcguto ErkrLrion; t.r.ral fcld 

pale, with a ddk blnd aloDg rhc supcdor na€ini Fdp@Dr, prca.di4 5 slighrly 

distant veint, Sc bearing 2-3 bl@h4, Wings very sno( 

BrowD. Dotrlcd wilh pa1. spoB on doBun, ycll*bh b€ne:tlr Cdci brosn. 

Ceniralh v.ry cl@ ro th6c of V. 6p.tt!,J (Fig. 4lC); .phhrllu wirh derio! 

megin d..ply .rd b@dly .turgiEre, pGl.rior nitobed, Dcdi.n lobc vcry stM1, wid.. 

lebly mth.d in 0E niddlc, reo lne..t tot6 r tid. preloqed; doparmG modified 

i.to p$crior lon8, hooL-likc attucrurcs, 

FEMALE: 

Il is siEild !o fuL, but diffe6 in rlE fouowing charcc6: 

Fae a lirtle @Mvc below forehead: chakr yc ow. Modibla notual. rheir 

yeuow @lour extcnding o. sid€s of lbe cttFu! up !o a.t nn t @kcts. prcnotun febty 

widcnins in {ion( latcr.l lobes for llte mo$ pan yellow, witt onty supcdor turgin 

turowly brown. Ligs and .lytra reddhh-brown. rhe lan r with a yellNirh humdl 

band, exEndilg !p !o O. 4t abdornjMl lcrgir.; dond fcld *irh f@bly obliq@, clos.ty, 

sp@d, r.guld rcitu, eaNrc.s rciolcE nundou!, fqnirg lorgi Mow caltri tacr.l 

field d.rl ir supgior h.ll sirh th. qi pal., Sc,boehcd t@ apex. (ripdiror tong, 

sraighr, wih apiM! valvcs ndow, larmlatc, snooth, poinrcd al apci. 

M€NNmotl (nD)r 5d 59 

E,L. 

H,F,L. 

6.D, R.!& Mtu S.D. 

12,013.5 12.73 0,49 

1.6-1.1 3,7J 0.19 

2.5.?.53 2 0.0t 

3.t7 '| o. | | 

4.2t4.5 a.53 0.13

0.56.0,62 

-t5G 

Holo.rp€: I d, l?'X-94, ,auhs$ad 6nushab) 

Alloqp.: I e. l4-x-94, J.uharabad (|(buhabl 

P.dryp6: JNhlabad 2 6, l+X-94,4 6 5 9. 17-X-94i Mith! Ttwe 

(Khshrb) I d 4 9, l+X-94;2 9, ItX_g. 

The spmin ns of tlis sp@i6 lttE fouhd in the lerf'lid.r undcr dc € fo6t 

pranratio$ of sufaida (Ercdt td c4tuUqlattb) .!d th8ehtu lDatbaeiq tk@)' 

Diltlrgohblng ch..r.t 6l 

This rcw s!€.ia! 3lpw3 a cloe Fldionstip *ith v. atltB6' fron *hic! it @n 

be s€parated by irs occiput, whEh ir liShl coloured, nandibt6 l*s snonglv lcngthcrcd 

and rhc genihlia wi$ m.dian lobe of epiPhallus verv 6hod, slighdv 16! produed a 

@nPf.d wnn y- 6P.64. 

Th. Mne of thh !P4i6 ha be€n d.rived fo,i its oeipul. which is li8!F 

Thc tyF dEriJ of $B ipdi6 hs b@D dcP@r.d i! rh. lDgr M@s 

Enlonology, Univ.rsity of Agicll$rc' Faislabld, Palntan' 

Dlparlnc of AgriculoEl 

MALE: 

v.tt Vi.tntut tu&.@tut s9.tur, 

(Figs. 424'C) 

12.0 mm), Body c.lounlioo brwn spofted wirh 

Siz snau (11.0 - 

vellowish

-157- 

DoBum dark breM. s wid. s or 3lidrly Mwd lha! prcnotun; @ip!t tiShr 

colour€d, prcvid.d wirh 5 bown ban&, rhc ou0er rqo uiird wirh u o6liq@ y.ltow ti*i 

forch.ad a lnde produced b.tqen dlentu€. btunids srdudly @iovc jusl bcloe lhe 

a enor cllu (Fig. 424)r f@ . linl. flaftrd. brosn, ic shicld Dryid.d wirh a 

n dian ycllow ba.d, .iendi.C up to anFrid @llus: cltT.cfonhl suture f.cbty 

angulnc, Eyes bl&k. Oc.lliy.llowhh, lhe laEral rwo unit d with a tnorveGe band ot 

lbe um @lour. Mandibl.s y€llowjsh, nuch shond (2 , 2.5 mm) than $ce of y. 

6p"6d, Ptlpi lighr bwn, th. luiu!.y wiih 4th s.gmcnr very short. 5I! .tongaE, 

wide.ing, otliqu€ly ruMte ar aFr. Ancn@ bown. widr lsr egmnr pale. 

Ponotun tldvm, witt sid6 slighdy bulging out in rhc diddl., eGrior 

Mgin fc.bl' 6Mvc, porc.id almdr 3raign! dilt wi! etun brown ald y.llowbhi 

lateral loba fd the ftosl pan bown, wirh a Mrw pale band along infcrid mdgin, 

wideuing arle iorly, cx[.ne i.fcrior Mein a@nding posLriorly, 

L4J pal? y.Uo*ish, nonled with browri a|lrlr iibi.E perf@lad on dErnal 

lace wi0 a ldg€, ovd tynptun, th. int rnal sitholt or with N vcry snall, rcund.d 

one: pot rior fcnora 3hon, s|oul, lris(d wnh brown, tne lati.r inrcruptd with rm 

longitudinal pale lircs or dlerBl surfa@; Fsreriq tibi& @cd qith 5 spin! o. eacn 

Mgin: pGariq bait$i with 6 i.r.nEl, 6 - ? cxl.mldeDrjcle. 

Elyra brcwn, .xt nding io lhc ap.x oflMohcni nimr alD6t $ long s wide, 

rou.ded posteriorly, dividcd at middle by a cwed vein, ill irte.nal margin Eror. frcn 

lsr chord (Fig. 424)i ih. larer unilld wi$ 6ir.or by a smU v.inlel n@ $e a .rior 

angl.. 2nd chord sEo.gly aB.d; h.rps sith 2 oblique veiB; .pic.l fEld 6dcr rhon, 

pr*ntin8 co.fused v.narioni lalchl ficld da.k. pt*nrine 4 - 5 dirhnt veitu, Sc b..rin8 

2 bBnchcs. WinSs shoft, 

B@n, noulcd *ith 'rllN on docuo, y.llwish blErh. C.ci brcwn. 

G€trihlia cxacdy simil& b thoF ot v. dpzttut, (Fig. 42C), wnh cpiphallus 

pGreriorly filobed, latral lobes a lidc ptolongcd, de nedio short, aln6r runote, 

wid a rnall oicd nolch.

-15& 

rl, ar. VWat r*-t i. -, tl.a-aF-d, 

hr.qr,S rrc'.tl-'-lid,

TEMALE: 

-159- 

Il is sinild ro thc b.lc q@pt thc following ch@t B: 

Face rcry littb nareledi ch.cls yellowish, a ddk brown band exlending b.!ow 

@h ey. up to thc mandibl6. Thc lartcr y.Uow, this dlour.xtcnding thr@gh sidd of 

cly!.us up io anicnnal sock€rs. Elym cxEnding !o tlE niddl. of aMoftn; do.el fEld 

b@n. with a ycllN huEnt bard, P@ning 4 f€. tcia, 2 bnEha of Cu, 

raBvce winlcb n!mc@s, forminS ift8ular, clonSrt! cclls, sc bding I ' 2 

branches. Oviposioor su_aight, with apiel valvcs @row, lan@laic, ss@th 

Msmment! (m)r sd 59 

Itolotyt : I d. I6-X-94, Midr Tawam 

Allotype I 9. l6-X-94, Mitha TaMna 

Ptoltpes: Jehelbad ((hBhab) 4 d 2 

Taw@ 5 d 2 9, l4-X-94;2 d, IGX-9I. 

Md. rdL 

n !3. Md S.D, 

ll.Gl2.o ll..O 0.37 

3.G3.5 ' 30 0.19 

l.rz.2 2,06 0.12 

t.2-1.7 t.42 0.m 

5.&1 .0 6.t2 0.t1 

l.tr.55 r.ll 0.02 

r.rl.9 r.76 0.10 

1.U1-9 7,r3 0.39 

2 4.2.9 2,62 0.17 

5.2-3.5 5,44 0.12 

0.59i 62 0.o 0.01 

2.95,3.13 r.06 0.10 

0.63.0.79 0?3 0.0.1 

0.7G0.?? 0,12 0.03 

(Khushab). 

,:' "j' 

',_31 0.21 

t.t2 0.05 

9. l4-X-94; I 9, l7-X-94r Mitha

,t@ 

All the sp@idens of this spei6 qerc @ll@t d frcn $. l@f-litter und6 deN 

foGt pldlario|r! of srtaita (Ercarpta Marlîs) etd sl@t6n (Datbqsia 

Dhlngx&nirg chr.(t rs: 

This spdics shows il. affinity !o v. .4o.16, Aon qhich it €n 6ily b. 

s.p@t d by it3 ninor which i. divid.d at l*t in siddlc, *ith int rnal Mgin EmE 

fiom $e lsr cho.d, agial ficld with onfused vcmtiotr, latcral field with 4 - 5 distincr 

vcjB and the posrriG tjbie *ith 5 spiB on @h nrgir, 

Th. md. of thii speie. h6 b@n derived ftod iB €lyrrl venation, i! vhich tllc 

ls chord is di!6r liom lh€ incrnal Fdg'n of rhe mirror 

nE ryp. mat dal of thh s?e.ies his ben depGicd itr thc lM:r MtMm, 

Depatme of ASrjcul o ral Entonology, U n ivers iiy of Agr icu lture, Failalabad, P.kkran, 

MAI,E: 

5. v.ttllictorsi .,ir,n!an sp. nov. 

(Figs, 4lA-C) 

Siz€ snall (ll.?0 mn). Body colomtion s[inins bl&k, wirt l.gs y.llNhh. 

Dosun shining black, slighrly widcr $an pronoomi @hur adorn d wit 6 

prenircnt reddi.h-yellow b.trd! ard e obns mcdid r.ddilh linc, O. @r.. ba.d on 

each sidc brcadly lnning dtlriorly qith $e posl! ior mrgii or €ach ey.i forelEad 

febly p@j@iiog b.ts€n dLnM., blek up io clyp@to.ral 3ur!.c, pr.s€nriq beE$ 

the d€rior @llu a lnall y.llow li.q ehdk! black, wirh . y.llov bard, the lalr.r 

.xterdinS fton thc orcr n rgin of eF ro rh. our.r ho!6 or tnrt'dibt|. (Fis. 434); hci 

..ddish-yellov: clyp@-fronral sutue roundly @Dvd, Ey6 bl&k. Ocelli yetlowilh. thc 

lare.al lwo unit d wilh ! nary, fdbly visibl., rEddirh-brcwn lirc, M.ndibb snqt, 

normal, Palpi broken. A.G'@ brown,

-161, 

Protrotun0,73 tins a long a wide, slightly wideDiDg i. f@nr, its 3id€salnosr 

parall.l, dlerior trwgir fehly @Mvq Dor4.ior rraight, both providdl wid brbrl*r 

dist rc{ly fl.t, alm6t glabrcB, brownhn-bhcl, wirh f.w t€llwisl spots, th. two ftd 

p6!.iior latenl an8l8 dore proniEnti laEral lotr.s wi6 suFrior pdr bl&k, rhe 

intirior wi6 a ycllow bard. broadening ant€iiody, ext€rne infeior nrsin blek, 

L4s ycllNish. prcvidcd with sp.e bwn spo6: dl.rir tibirc wirh rymparun 

on both sid6, lhe ouer, laige, oval. fie in@r, v.iy snall, on.ded; posrdior fcmon 

thick, ob8t, 0.38 ri66 a vide a! long, $rhred whh brow., obliqu. lins otr t.ir 

cxtcrnal fa6; posictior tibbe dd |!rsi brown, t. fomer dmcd with 5 i erMl, 5 

ercr..l spiB, 3uPcrcintermls {rd m.dioinrern ls .qual, 0.6? tin6 6 long s 

bdilaBus, the lalt r prdi&d witi 5 - 6 interdl, ? crcrnd de iclc.r, 

Elyka brovn (brok€n at apex) i nircr dividcd bchind middl€r lsr chord esuhrly 

coNcd, 2nd sro€ly situaE, 3rd alnGl st aighr, lsr !ni.d with niror by a singlc 

v€it'ld n@ anEdor an8lei hdps *ith 2 mdly snight vciB: apiel fi.ld vitl confus.d 

v.natjon (Fig- 434)i l.tenl fi€ld bl&k in sup€rior half, th. Bt liSht brwo, provided 

wilh 4 veiB. Widgr very slron, 

BlwnLh-bl&k on ddsuo, brcwnbh-t€Uow b.EA. Cqci brck.i. G.nimli. 

(wh.n viewed Aon above) very d* to thoe ol n aly@r6 sp. mv. (Fig.43c), 

dxcept the pcrerior m.lian lob€, which is long, M.ow, slightly eoarsina0c at ap.x a"d 

antc.io. cndgiEtior enh a 3n ll rcdid michi eiop'merd wry rinild io those 

Mearurcme.ts (m): I d 

P.W. M.L, H.f.L. H,F,W. tt,t.L. 

1.0 l-9 7.9 r.0 t.5 

E,L./P,L, M,L,/M,Itr. H,T,L./H.F.L

It! .3. 

I 

-t5z- 

u.a.W tu+t Ab.dd;ra, 

h-dt l,{tn drc'iF-r tH*.

H.bir.r: 

- t63, 

EoloF!.; I d. l5-vll-93, Ninbdl (M@). 

A single malc wa @llfEd frcn gres lndcr a pine planftion. 

Dbtl4uitinr chulctes: 

It ba r d@ Ilsbla@ to V, c.!L,n

-164- 

Fk 4. V.lattt utu lala .eld.ll4 dd-l.t d.

MALE: 

Mhn*ali I 6, +X-8E. 

,t65- 

8,L.1?,L. 

A rj.el. hrL h,s b€n @llelcd iion liSnttrda@lriEredfi.ld. 

This specid ha betr cor

-155- 

Elytu @mpler.ly dek b@pn, shining. e&nding ro thc rip of abdohcn: niror 

loz€nge-shap.d, wnb rcundcd pGterior angle, divid.d nuch b.hind middlc by a cury.d 

vcin (Fig.45B)i diagooal vein very close to the ftsr cho.d; harps with 2 obliqle veitu; 

aPiol field v.ry 3hon, pr.scnting only 2 v.iN and I wide, ndty irEgutd rdicularion: 

latcral field d.rk brown, tnnslucenr, wirb 5 regularlyibi. v.ins, Sc bdi.g one 

Brcwn, wi$ cerci ycllowish-brown. Epiphattus wnb ncdiad tob. having a 

shallow. almost roundcd noich, ia latcr.l lob* shdr, blundy poinrcd (oedi.n mrh and 

lac6l lobca slighdy rhorld rhd those of y. /d{4r)i dr.?.tn'n.c, loi8, i.row, 

lr@ked (Figs.45CDE). 

M6umdtr (tm)r I d 

B,r, H,W_ P.L. P,w' 

|l0 3,4J 1.30 3.r0 

E,L, M.L. M.W, H,F,L, H.F,W, H,T.L. 

t.ro 2.o 7.0 2.7 5.0 

H.W,/!.W p.L,/p.W. E.L,/P.L. M,L,,nt,W. H,T,L_II.I.L 

r.0r 3.33 

Eolotyps I d,9,V111,88,R.walpidi, 

A sinelc male was @ll@red at tjShl nd . deNe vegcilrio. along rold ride. 

Dlningunhhg chrn t€B: 

It ha a very close rcsemblane |o y. tuar, bd @. ealily be epararcd by the 

br@d had. divided nnd, wnolly ddl brcw. lalcral fi€ld, wilh 5 r.Suldly,Aaed 

vcins ad the g.nitrl @nf8lntion. Th. larcr aho rhows ir! atfiniry lo rhar of y. 

lattl'd. fron which ir difi€c io rhc @iput, qhich is providcd wilh ycltow tiB, 

whitEh palpr, dym sllh 2 h$p verE and much 3horlq wrngs. 

saed (r9E9) enomouly ideotili.d ihis spai.s a S.aptipedd lallar. br on 

th. bdie of dE ahore diff.E'B 't 3h@ld b€ rcgddcd 6 a *w spei6,

tu.4s. 

B 

-167- 

V.b,{wk'Nt dlridu lp. u. A, h-'l rd !.!nodu d, 

blrd 4.ci I, drr'. ar C-l' F|l$ d; C, .tbt tu, rlor.r 

.a.dJ D, F, phr. Mdd, r.d.l rd ltE l .{.4. 

c

-t6E 

Thc name of this sp@i.s hd ben derirrd from its divided minor. 

Thc holotypc of rhis spei.s hs ben de96ir.d in th. Iiser MEun, D€pstnrcnt 

oi Aericuhural Entomologr, Univeuily of Agricuhur., Fai6.labad. P.kiltan. 

C.du il4od,.otrrrra C[op{rd 

1951. Gry4rr Chopard, Re.. S. Ausral. Mus., 9: 1108l 

l. Md[Eott llla C[opsrd, E6, Madr.,37:2?2. 

Tt?e CrylrB @dp.d6 Scha!6 

The cou€.ted maErial cnrirely agrc6 wi$ O. publilh.d daription (otc ad 

C.de, 1984b) of thi! gcnB ard Ls rcpEsnrcd by 6 3F.i6 in rhis rcgion. 

t. 

Ket to rb. $.{i6 ol todi4rsrt tur CboD.rd 

witho[ a yellowilh bard b!tw@ latlal @lli; genicali. wirh ! h.di 

2 

bad b€r@n la{.r.| Gllii gcnibji, wirho a mdtu 

2, Sv. small /lds dDn 10.5 nml: h.ad wirhout any mdking: .piphaltur s i a 

J. 

(Fig . 468). 

(Fis . 4EC). 

small, roudcd menian lob. (F'g,468), 

Sir ldge (gpt r than 13.0 mm)i head enh thre bmd brow! b.rd! on 

dciputi epiphalls witl a wide bedid lob€ (Fie.478). sisriarorr (wrDc.) 

Epiphallus wilh pGErior .@ginarion U{haFd (Fis. 48C,. 

Epiph.llB wid pckrior md!'Mtion V.rh.ped (Fig. 50C,. 

Occipu( br@dly dng€d wilh yelloq, wi$out .ny rukingi elytn wnh hteral field 

palc, provid€d wirt tEly sFaighr bur fdbly-spccrd vciN (Fis. 4EB): .piphrlta 

wi$ posictior emrSimrion deeply U shap.d, withour . oedor 

mrch (Fic.48'c). ,.....ntde (cboD.rd) 

5

-t69- 

Occipul black, wiih 6 short, ycllowbh lin6i .lytra wi$ latedl fGld snol(y. 

p@ided with sinuate, sidelyipac€d vciN (Fig. 49C); ep'phallu *ith pocr.rior 

*T:lll:1 " l* ::::1 ::* " :*:l*:il,..- 

5. Body srawolo!rcd; clyFcfron|ll 3uturc ltrongly angulate (Fig.50A), @ching 

almd clN !o fic aot.rior @ll6t cpiphallus wi$ posrerid endgiotion d..ply 

and finery v{har.d (Fig. 50C). .. . . . . . . . . . . . - . . ..ot llu sp. Nr, 

_. Body ,rllNish-brownr clypco-fontll sutuF slighlly mgulale or I@ly stoiSht 

(Fis. 5lA)r epiphauus with posl€tior €maryiMdoD shlllow and broadly 

vihaped (Fie.5lD). . ,ttutuquit sp, noa, 

r. ModbogryUs ,Lrtrr (SrM) 

(Fi$ 46 -D) 

r9n. G,tvadd ,/!,M S.B!rc, Mcn. Soc. C.evq25:215. 

1928- A,t at fl@ipa Cho?td, Spol. Zyl., 14: 197. 

1931. Grr\uls ,/..&r ChQatd, Bull. Rrfll4 Mus., 6; Iro 

1961. Modj@gry\tu ,lat[t Chopdd, Es, Madr'. ]7:2?4. 

1969. Gryltut bL46 Chopdd, FauM of lndia, crylloid., p 55 

It gr€dy lallies qi$ dE plblishcd dcscription (Chot&d, 1969i Sa€ed 1989) of 

ihis sp.ci6 ex@pt lhe p$tcrior tibiac which are wih 4 - 5 interml, 5 external spr6€s 

mirror 1.0? tin6 s long s widc (Fi8- 46A) a.d elvtta wilh lateral field in femlc 

pEsening veiN s dbiant 6 in nal.. Ccnii.lia (Figs 46BCD). 

MesuM.nts (|m): ld 39 

M.|. 

s.D,

Ela.6. 

E 

!tudt@grUt, thNB, A, drlr d, eD, rdi.rb di 

a, edlt llt4 do.-l !p.dJ C, D, phllc ooda, qarl rad

.171- 

,.;., 

t.3-1.35 

'; 

r.33 

Faisalab.rl 3 9, l5lx-E?r I d, urlabeu.d. l3Ln fon rhc srud.trB @[@rion al 

rh. DAa nenr of Agricultuhl E.tornology, UniveBiry of Agricutturc, Faisatabad, 

All thc sp€ciheN of rhis sp€cis were @llcFd ar tighr F{r cuhivar.d fElds. 

2. Motti.osrua slr{tra (wittq) 

(Fir3. 47A-D) 

1869. Grtllu sisnil6 walk r, car. D! u9r. Sal6r. Brir. Mu., tr 38. 

t961, ModieSryll$ ri,rir,@ Choged, E6, Madr,, 37:2?4. 

The @lldt4d nderi.l agrs wid th€ publish.d d*iiption (wattd, t869r 

Chopdd, l%9) of rhis sp@iB, but difien in ttc fouowins cnan as: 

Prom$m 0,59 - 0.53 rimes a tong a wide, stighrly @rcwiig ii frcnr, wi$ 

anbior MSin f.rbly @Mvci dnk witn 2 l&!c hl4El 3poB: pcEior ribi& emcd 

*itb 5 inrernal, 6 ext r.al3pin6t elyua id male wih lst chord uniritrg rb. mir.or by 

,;

I - 2 @id.c (Fi8, 4?A), 

cciitaria (Fi8s.4?BcD) 

M.6lrtnn.s {na): 3 d 

Sc 

-tn- 

brring I - 3 bleh6, *in$ p.cscrr d 

2,7t1,4 

2.2! 0.r2 

3.13 0,28 

Abbo6!.rl I d, ll-vll,93; 2 d, untilcltcd, Irl@n aoD dE sflttclB' @th.rion 

n rhc Dcpdtneni of Acriculhtrat Enlomlogy, UtrivcrNiry of Atri@b[., fa!.t.brd. 

A rii8le illc 3!c!imn h!. b@ coll@cd &on lighr rc& ! ithL foBt 

h n s b@r @Ucci.d fa rt fur ri@ filE Abbod.b.d. E rti.r, ChA$d (t96p) 

@oded it I'om Lydlpu (Fai!alab!d). 

Thc sp@iocB ollacd n hiSh aldndc in mourr.im hrvc nor. d..k g.n.hl 

@lolrataon, with pronoral m&kins! hddly viiibl..

nC. 4?. 

-1?3- 

rrdrilt&ruat dgnito8. a,.rin d' FD, idrdb dr 

B, Cd'rllutr 'l6d rF.d' C, D, p6.llh mDra, Errd ud

-t74- 

3, Molli@muus .bMi (Cbopord) 

(Fisr. 48A-E) 

t%1. Crytta .hsad Chopad, E6, Madr.,37:273. 

1967, Modiaeryll8 zAtdui Cnopdd, O !. C*, pes 10, Grrll., p. 81. 

The collcckd mlErial of dri! sp.ci4 csembla rhc published dcsciiption 

(Choprd, l%9) of tht sp@i6, bur diff6 in tn followins charclen: 

Proiotlm 0.58 - 0.63 tir6 6 lone 6 wide; pci.!ia{ tibia. with 5 i !rol,5 - 6 

ert rnal spi6; nal. clyra a lorg d or a lildc shond dd abdoneni nircr dividcd 

a hdc b€hind niddlc (Fis. 4EA), Sc bding 2 branchs (ftrc1y o@)i fenale ctytra wirh 

lat .al ficld 6 in milc, Cennalia (Figs. 4ECDE). 

M.suttmnis(m):4d 39 

s,D. 

3.ltt.5 

,;, 

a;

F1!.4t. 

-175- 

_MltLot4lu .h41 A.8\drh. d,,ro,l ud Lt r i*r{ 

e{5 FL5 d; q .rtLk 'rrt {.4 D, r, lt.|rt 

tu0.a, rll.l .rd Lnrd -6.

-l1G 

FaqiBbad (Kohat) 3 d 3 9. 3-vlrl-93; Rawalpindi I d. ?-Vlll88. 

The spccincu of thk spei6 qft found in 

ud* dense plindions alor€ rcad sid*. 

It ha b@n @llcrcd for dE fisr ih. Aom F4inbad and Rrw.lpindi. E5li.r, 

Chopad (1969) descdbed ir rrom Maln (Karachi). 

MALE: 

Herd: 

4. Modi@s,rllat b@cut6 3P. Nr. 

(Figs 49A_n 

rhc rcil c@l! and on the lafli&r 

Sia d.dium {13.0 - 16-0 nm). Body bl&t. itP.ring pcEriorlv. 

DoBum blek, Mlyss wide a! Fonotumi occiPut &n€d with 5vellowttripa, 

de ol.r rwo unir.d *ith an obliqE v.in; h@ bl&killr in sup.rior pan. b@nilh on 

ctypeus, labrun, and mandibl4i clp@Jonitl 3u0n .lm6l straight (Fis 49A) Evcs 

black. Oelli ylllow, Iar.ral two mnn4t d wilh a @nspicuou stiPe. Mdiuarv PNlpi 

ycllowisn, 4th sgnmt short r tha 3d a.d 5d $. latEr darLn d, s'd obliqucly 

ruMte ar aper. Antennac brcwn. 

Pronolln distinctly br@der tMt long. 0 54 - 0 60 tims d long 6 broad, sli8htlv 

widenins in frenr. itt anffior 'ugin rcry teblv 6Mrc' pd!'rior staighl both 

p.ovid€d with dlN ro*s of brisri.si dist SeMllv dart bul febly par'rn'd with 

ycllo* and da* spots in $c posterior balf; lareBl lobee bhck wiil 'ntctior halfhlving 

a na(ow ycllowbh ba(l !t cxrrem infcrior mdgin. eF.dna rcdued to a sMll psre 

Llgs y.llowish, molthd witn a fcw dek spotsi anErior tibia. p'rforated w'$ a 

larB., owl oulct, and a sFrll rolnd€d itner tympeum; pdlerior r'non 

0.91 - LO5 rims 6 lo.g s.lysa, with 3ttong ddk oblique strie on bdh sids, whidr 

arc inteiiuFed qith rso btoqtrish_tclloq loDgitudinal lin6r posrcrior tibiae 0 66 - 0 80

-tn- 

tid.s .e long s f.no6. dmcd rnh 5 i erMl, 6 cxt nal spinBi superoi lrol and 

medid apiml spuB alo6t equal, superexternals shoner lhan nedids. 

Elyua cxLndinS alMr to tnc aIEx of abdo&f, or a lndc ihon r, 3.24 - 4. l0 

times s long a prcnotum, dbtjrctly ljp.ri.g post€riody, d&k brcwn pith lighr hun ral 

bands, almct uapdni dino. longcr rle side or rcry @l' a loig a wid.. with 

rounded pdtErior angl., dividcd in rh. diddle or sliSltly b.hind by a curyed vcini 

chor& acbly b.nq h.rF wirh 2 obliq@ v€iai apiql fcld wirh 3 4 e.ia. rdiculalion 

formed of alEot square, rery Egdd calls (Fig. 49B)i lsteel lield snokey, tllBpenr, 

wilh 5 d2rk. eid.ly*t.c.d EiB, S. wirh I - 2 bl4h6 (Fig. 49C). Wingr @dra, 

Blek abdr, ycllNirh bcHtt. C.ri br'Milh lo bldl. Cditrlia 

(Fiss. 49DEF), wnh epiphallus H shared, havine a shallo* anterior, and widely urhated 

p6r.rior cMSiedon, $e larl.r pnid.d wirh r fG.Uy vilibt. dEdie mEhi 

api6 of lhe posbro-heral prcjetioB dd 6ropdmd6 cw€d upward, the latEr, 

ldB., .rending beyord rte hi.rd lots ri vilibL in rl'. doBal dpel. 

FEMALE: 

k is sioil& !o male. exc.pr UE follwin8 char..Ers: 

Porctln wft sliddy sinul! pGt rior @gin. ElyEa a long d d stishdy 

longd thd fcnora, 3.53-3.78 tir!6 al l@B r! po.on; dd$l fEld wirh 3 fi4 vciB, 2 

brarch6 of Cu, crc! vciB nuherou, fqnine rcguL., alnoll s4u@ a6t&i lrr.ral 

ficld wnh 4 *id.ly diitaot veiil. Wiirs small or dudare. Ovipsnor boc (2.01 ,2.15 

uds 4 lonc a clytra), sEaighr, in.ok!l v.lv6 @r@, tamatatli rcur. at aD.x. 

MeauM@t! (m)t 5d 59 

a.L. 

s.D. s,D, 

,:] 

"::'

5,2.6 2 

r13 

,178- 

I, 

P.nttp6: klamaM 7 d 4 9, lt-tx,94i AbboroM 2 ?. l4tx-94; 

Faisalrbad I ?, lGvll8S: i d, r6-VI q3j Coja Oo6a Tck Singh) I 9, 29IX-EE: 

Mimwali 3 d 2 9, 4-X-6Ei Patpaun I d, l2-X-E8; Rljanp( I d. 2t VII,88: 

Rawalpindi 19, l2,Vltl88; I d, I6-VIII-E8; I 9,2GVIII-88; I d, l_tx-E8r I d, 

3-IX-EE;3 d 2 9, untab.ttcd, rakcn Lom rh. stod.n$, oltecrion at rlE D.?.rrlr.nr ot 

ABricuh!61 Enlomolo8y, UnjveGity of Asricur$E. FaiJalrhad, palill.rr. 

This sFci6 qa! @nnonly folod in fie leaflnEr undcr dene iorcst ptdiatioB, 

a w€I a on gBs undd liShr stues. 

Di*i.la&hlry ch.!.c.6; 

rk $dta Modi@eryUo !i46, itr Chopud (t%9), bur diff6 fioft rn 

published ferDle in h.ving (he @ipur with 6 y€Iow sr.ipe, a @row band or $e se 

corour beNceo latcral oc€|i ..d the piorcral dhk wtjch k dndr d.rk brown in its 

k is are very de.o M. tutu is inqtrgu|Ace (l9E4b), from wnich n 

@n 6rly bc sep.ral!

nt. '19. 

oo 

o'o 

-179- 

tlo.U.o8r]Jlllts bonculu tc.n I, 16 ar & C, dtn d, 

tla-l rd l{.|| ndb DrF! dL d; D' Crt rs tb.rl 

..0..! E, r! drn dnpL*, rei.nd @d .{..t'.

,180- 

Tfiis spdi* shows a stighr variation in body cotour. Som. spccimeN cxhibir 

erclGiv.ly blak @louarion, whit. thc olhcB ar. dighrly lighE. 

Sae.d (1989) pla@! sne speimeB hd.' Crylra Jisrpd duerotheirctw 

r*mbl.Jr€ with rhe lakr, B he abole difleE.cg show rhal t.y &rualy belong to 

The @Dc of rhir spei* hd her d.riwd fren irs occiput. which is provided 

Ti€ rttr nar.rial hs ben dcDditcd i. rh€ In@t Muslm, Depann€nr of 

Ag.iculrunl Enromology, Uriv.Bity of Ardcut$E, F.ietabad, prtijhn. 

MALE: 

5, MolLomUB cordllB sp. aor, 

(Figs. 504-E) 

SiE shdl {t2.0 B.j mDl. Aody s@w{dourcd. 

Ycll*ish, slighrly namer lhEn p,orcllo, @vded. w'rh v.ry Rre pub@: 

o@rplr sith 3 broad inkr and 2 mrrow out.r, shon, longitudinat brcwn bads; v.rEx 

w'$ a br@d, ransv€ac, broen band b.rw€e. eyes, a.d a s@nd onc of the sme cotolr 

otr lhc fro..al coun.bov€ oed j.n @lt$i froot l crrun febly swottcn b.r*@n rhe 

.nlen@; r@ wirh p6rctypcus brown. anrctr?cu, ,c *, Irbtud .rd mrdibte p,r. 

bDwn: clyp€cfronlal slrue srrongly angutal., r@hi.g very cros. ro ancno, oclllls 

(Fig. 50A), Eyes bt@tish-brown, Mdi ey palpi ye ow, vilh 5rh seghert obliqucly 

rrunerd, brown ar ap.x, An€n@ y.uowBh. 

Pronoom 0.57 - 0.5E rjnB d long s widc, v.ry fably widc.ira in iionl. ir 

atrFrior ntrgin @nqvei poterior straightr disk atndr l!r, yeltow, wirt a nedian

-l6l- 

longitudiMl brcwn line and 3 laise bioqn 3pots on @h lide alo.s mdid linei tareial 

lob.s brown, inf.ric D&gin eirh a !a!lw y.llry ba!d, widening rowrd! aar.rlr 

angle, the lauer almoskound.d, .xrrenc infe.ior f,dgin snonSly.e@nding backwards. 

t4s yeuowi ellrior dd n diu fen@ febly spored qirh bwni &arior 

tibiae FrlotaEd enh a l&ge, oval olrer and a small, iound.d in@r tympuunr pdErior 

timoa 0.94 - 0.99 tim6 e long s clyrta, yellow. v€ry ligttly holrld wio b.own; 

ponerior tibi& 0,68 - 0.69 tih$ s lo.g s pcr..in fcm6, dftd win 4 - 5 inrcrDal, 

5 external spiGi supftint .nal apiol spuB sliehtly loiger r'ro mcdiaN, 

hcdi@xarnds long.r ri,n o$cB, 

Elytra slightly shorEr thd abdomen, 3-60 - 3.?9 tiines s long 6 promrum, pale 

brcwn, t aBh@nt; nim 0.94 - 0.9? rin6 a long d widc, l@rtcihrp€d, rcund.d 

pdtetiorly, divided by a curyed vein slightly bclow rhe niddlc (Fig. 508)i diagonal vein 

aln6t st aighr; hdp! vith 2 obliqrc vciBi cndds fably cuNed, n@ly pdlu.lr .pidl 

fi.ld ralhfi shori, pr4nting 3 v.iN, reticulation fomed of v.ry @guld, elon8.r. @llsi 

l.mal field p.le, e,i! a brcwn ba'd ir iuperio. p&t, pr*ni.g 4 fcebly dblatrt veiN, 

Sc witn I - 2 br@hB, Win$ long, eudlE. 

Shi.ing bl&k lbovc, ycllw bcn aln. C.rci ydlry. Subgcnihl pba b@n; 

epipballus with basal portion sliShtly produced dorso-poslliiorly, ulerior Mgi. broddly 

blt eeakly cMgidrc, pocleriq v€ry dccp ard rdrdly V,shap€d, la|.nl lobee very 

naiiow, shaaly-poinocd, upwardly{urycdr 4loparmeB loo8, biilrqrc ar ttEir bM, 

rhon r dm lgterd lobB iD do .l sper (Figs. 50CDE). 

FEMALE: 

It is rimilr 0o the malc cxept rh. following chdui6: 

Prcnotln with laEral lobB ycllor, prcvid.d with ! mow b@wn band in iB 

upper pdt, Elytra 3.21 - 3.63 rima d long s pronotum, pale, wi$ rcguld obliqu. 

vej6, F.nsErs v.inleE luncreus, foming long, namw, almr regul& @llr, 

Olipositor l 02 - l 19 tims s long 4 poslerioi fcnorai straight, irs apical valvcs raihe. 

sfton, wide, @ur. {r atcx-

Flg. $. 

D 

Ndlatltll4t @.dtttt r. o. ^, n . a; a, .trn a; 

C.r- rdr.r| d; c, CrEft, &rl r{.d, D, E, AIi. 

onDld, rdd !d La-rr ar.dt.

Malllldr!(m):5d 59 

Mn. 

-t83- 

3.051.5 3.2t 0.13 3.t-3.J 3.29 O.t3 

IloidyF. I d, lcyll-88, Frieht d. 

Alorype I 9, l6.vn-88, f.ietrbad. 

Prr..tF. Krlrr I d, Gtx-93r Frj.rt.lad 5 d t? 9, t6vtt{8: Cojn (To6a 

Tek Singh) I d, 2ElX"8Ei Muaffargdh I d 2 9,2GVlt,8E. 

Hrb|!ll 

A1l lhc ap@in.Ds of this sp€cie, tle b.€n 6tte0cd froft lighr ncir cllrivlr€d 

6eld5 .d along r@d sid6. 

Dlrthguirhlnt chs.d..sr 

h is vrry crd. lo nd@{tnu Uwqt in Chop.rd (t969), bsr 6 d , 

,; 

".,,

-184- 

bc scpdr.d by i6 hed patr.rn, pon€rio. feoora wnhour b@wn sniatioB, mate.lyr6 

wi6 lpicil field prcscnring 3 veiD aDd a quir. differe male gcniraj smcturcs. tt atso 

shoB ia close @nblan . @ M- bwdigatc8it, bur ,lighrty did.B in r!. ,nale 

gcnit lir fig!rcd by corcldov (t98) io havi'g the Aiphr u eirh poslcrior 

cm.reiMrion reu'dly Vihap.d, tn El lob6 wirh oucr sid.s .tndt sEeigh bclw rh. 

level of post€rior .Mginarion, ditrercne in eioparan B and lateEt view of the 

cpiphallc (Fis- 50CDE). 

Sa.cd (19E9) pl&ql a nmb.r of rFdreN of rhir lpccjcs ni Gryllus 

lt .,iapr oi rhe bais of rhcir .tosc rsnbllEc ro ric l.n r, bur rhc abovc difLrc'B 

show $ar they !.ru|lly belonS b tht n€w sFci6_ 

De.iYrtioo of |raml 

The nde of this species ha b@n derivcd f.on thc swo .D fronol ,Ntium 

Thc type Micri"l of this speiB hd b..n d.poli€d in rtE Ller Meun, 

DcP3nft of Agriculru.al Eotonology, Uniwciry of ASricutrurc, Faietabad, patishn. 

MALE: 

6. Mo.li@g,tutt qtn titpsit .0. w. 

(Frgs.5lA'D 

snall speci* (9.2 - t2.0 mn). Body colouradon deK Drown_ 

Shining ddk brcvn, nedly a! widc s pronotuhr occipur lighr colowed, with 4 

rably vnibb r€llow tin6, rh. ladlr @tou al$ drediry behind cy6. a n.mw band 

or lhc en colou. dr.rdilg b.t@n larcmt @llir fae llighrly widfi 1hnn long, brwn 

i. suFrior pan, wirh r y.llw tine bcpdr nr.dirn o..Us, yeuowilh o,r clr?€ls and 

nourhpaisi chek ycltowi cty!@ftoni.l slorc a..!ly aigutaL (Fig, 5tA), Maxiltary 

palpi y€llowish, sEcrines tighr brown in ddk r sp.cime6, Sd join dek ne

,IEJ. 

Promtub 0.59 - 0,58 tine3 u loig a {ide, iB antlrior nrgin r4bly co@ve, 

,osr.rior alm6t strlidt, sid4 sliddy bulsils ou! dirk n rrly nat, y.ll@ish, witn ddt 

brown paEh6. tlE l.tt r nuch norc dcvclopcd in ddlG rp@ibeN, pdid.d witlr 

squeF! sndl brisll.li l.t hl loh6 blackish-brNn, wirh a E@w yellow band alone 

inf.rior mdgin ert nding $tEriorty, iB .ri!r@ inferiq mgh bl&k, rlighdy s@nding 

t gs yellowth, notdcd wiih browni anreior ribi.. p€rfoared with a Lrg., oval 

qr.hal, sid a sn ll, rounded inlerial tympanun; p6t do! fcmola slishdy tut ned at 

ap.x, 0.91 - 1.02 dncs d loig d elyra, prcvi{kn wnh dark, oblique sri& on cxtctul 

fM, thc lan6 in&trupGd by 2 ycllowilh longi$diml litcti p6terior tibha 0.69 - 0.74 

rins as long d f.nota, prcvided eith 5 i .rn l, 6 dt rDal spiB, 

Eltt r s lorg 6 o. 3li8!tly shod.r thd atdonen, 3.2? 4,05 tiG a lory d 

Drcnotun, daik brwo, with light humcRlbdds: bircr lon84 thm b@4 l-15 - 1.35 

rimes as lons s br@d, tounded p6tcriorly, divided behind middl€ bv a curv.d vein 

(Fig. 518)i ddgoml v.in alMt staiSlttr chords tseblv cutued ndlv panll.!: hlps 

wirh 2 obtiqE wis; api6l fEld prdid.d wi! 2 I vei^t rcdolrtDn fortEd of !@lv 

sq@rc, cguld @Us: l.r.ral field traBpenl pale *itn bro{n band in sup'rior part. 

5 reglldly but nodedely{Pacc

t" 5r, 

o0 

.1EC 

Ud6E flat d@lt ?. d. A no dt 4 .lth 6l 

C, db&'l'tt Dl,ae d; DrdL!6 darrFr 

E, f. 't.[r. o@lB, rdd .d bi r4rdr.

M6urEerts(ml.5d 59 

2.5-1.2 

-t47. 

r.*.t ,',, 0,, 

Holotype I d, 2l-VIIt 94, Had klam (Vchei). 

Allo.yDcr | 9. 2l-vlll 94, Had kld (V.hdi). 

P..dyp..: Pshawd I d. 3l-VII-93i 2 d 2 I, l-VIIt-g3; Hqd Balloki 

(K6ur) I d, l5-vl-93; F!l!al.b&l I d, l2-vl-88;2 d 2 9, IGVII-8E; coj6 (Toba Tck 

Sinsb) 2 d, 28Jx'8Ei 3 d 3 9, 29-lx-88i Khi@wal 3 d 14 9, l8-vtt-8E; 1 9, 

rg-vfr-EE; Mi..Mli 2 6 2 9, +X-88i Mirha Tag@ (KljNlta!) I d, t4x,94i 

MuaffarSdh 9 d 19 9, 20-vll"88; Pakpatrao I 9, 16-VI-6E| E d 34 9, 8-vlt-E8;5 6 

l? 9.g-Vll-EE; ll d 16 9, lGVl!-EEi I 9, l+Vll-88: I d I ?,25-Vtt-88: Hcll 

PanjMd (M@fi&8dh) I d 2 9, rx-93r R{jaDpu 7 d 6 9. 21-vtt-88i Rawalpindi 

I d,9-VIII-88; I d, lGvlll-88; I d, l5-Vlll-EEi I d,25-VII,88: S&godha 3 d, 

26-V-88; Hydcrabad I d I 9, 22-X-93. 

Htbirrr: 

Th€ sp.cineB of |his lp.ci€s Mre l88ely @llccled irom gr.sss undcr lieht

-188, 

soures ntu cultivabd fields, rhe leaf-lirter lnd€r deisc foel planhtioN and the gr.ses 

DinhgulhinS cluncr.6: 

ft hA a cl* rarnbla@ !' nr'. t$,bbtun t, bu si8nifi@tty diff.B in Mtc 

glnirlia {hich dc al@t 3init& ro rh* of ry. tietqpr, fren whi.ji ir difrcB in 

clyp.o-fro rl surorc aloGr lF.ight or rcry febty angllar.. Ir is als v.ry cl*ly 

rcta'ad $ M. brchdXa,botdiffeB jn rhe mate Seni|atia figwcd by corokhov (t9?8) 

in which the eropmmc.A rrc fircty.h iahr.d on inrlrn l m&gi. bur cnding in . 

dightly h@ked, iow{dly crN.d apex, 

Sre.d (1969) pla..d some speim.ns ofrhis speies urdc. G' tniobscuN dnc 

to ihcir close r€€nbl6@ ro rhe laner blt $e dkstion of rh.ir nd€ g.niblia durin8 

ft. pranr s$di6 reval.d rhcn to b. a icw spei.s 

Some sF.im.ns or rhb sp€ci6 show almdr a bl&k Sencnt @ldntion ad rhB 

look Inore stronSly stoited whh brown on head, pronotun and lcgs. 

Thb sp.ci6 hd bai med h.c:ue of i6.nrirctt ditf.rcnr natc g.nii.li. ftoh 

t\os. of M. smiobtaw, wi$ which ir hs closc appeefl r.sembta@. 

Tte tyF nanrid ha t€en dcp6iEd in rh. t'gt Mu.un, D.pan@nl of 

ASricultural EntonoloSy, Univesny of Agricuxurc, Faisalabad, Pakilta.. 

Subfamily Nenobiinae 

Th@ spaid undcr two genera have b€en identif:ed in this subfanity, 

K.y ro tlE generr of N.oobiiDd 

| . Pcterio. f.non uaifornly pale-brown, wirhout ban&. 

P6terior fcnora y.llowish, with 3 rav€r6c bl&t bands 

(Fig. 52C).

-t89' 

c6ts nnn tuUu J@bso a Bt.ictl 

t8r''5. Netub l' Fi€bd, Abhandl. Sohn. ccs., (5)3:40a. 

l9c4. Pt@a.tubiw ,..obsi & Bimhi, Pr.n.i, L@hn, Rs. Imp., p.450. 

1913. BMbn Mbiu H.bdd, P@, Acad, .ar, Sc. Philad-, p. 443. 

Type: Nenpbi6 r.to.d Saussure 

Thc olldted nat .ial of . single sp€cies cuctly agB wi$ plblish€d 

deripdoi (Chopdd, 1969i otE and Al.xari€r, l9E3i sdd, 1989) of rhb 8.rs. 

r. Pt@a.tubiat @n@b. (wdto, 

(Fier. SBDE) 

t853. GryUu! hzydeni(1. btuchtpr.fd) Fischcr, Orrh. Europ., p. lE5. 

1455. N.tubiB lor.el,i C6.a, FAud Nap., Od. cduid., p, 37. 

1E57. N.tubis he!d.4i lt. b@htpt..;) YeBin, And. Soc. cnr. Fr., (3)5: ?61. 

1811. En.opt m .amlor Wdks, Cat. De.napt, Salhr Brit. Mus, Suppt,5, p. t0. 

la . NtuUB rarrt4r S.BuE, F.dsbento Voy. Tut6hD, Onh., p. 36. 

tEn. N@bia .?lori.(J SaBs!rc, Men. Sa. C.Eve, 25: E6. 

t8n. PEren Mbi6 ld.mtis Sauurc, Iden. Ibid.,25: EL 

l89a. N.tubi6 rerlrri Bun. Trd. en1. Sc. Lord.. p. 52. 

l9o4. Ptercn tubi8 rat@ Jrcober & Bi4ti, Prcm.i. tozhf,. RN. 

lyb. C)Edlph4 @n rlrr Kirby, Syron. Cat. O.th-,2: 81. 

1906. Pt4ontubi6 .rrloricu (irby, IdeD. lbid., 2: 16. 

l9o9. N.@bi6 hat.Lni wt. rheMa6 (nu$, Dbch, ent. A!chr,, p,l, 

1916. Ptron tubiB llztlc4i (t- bt@hl\erc) CigliGTc, R.dia, t2:286. 

1917. EaoontuUB la.nlb rt. tlt.M Z.chd. c€ndfllg_ Drs.hl,. p, 244. 

1911. NmobiB hnaw w, tch.lkowikovi Sbchelkabo-vtz.v, Bull, MB. 

1922. Pkrdtubiw eii.ntti B.rland & Ch@dd, Bull. MB. Hi5r, nar. Pri!, p. 235_ 

1y26. Pt.rcntubia httd.ni cotolot Uvarov, R.v, R. Enr, 20: 153.

l9G 

t926. N.mbiu ktt nlit rt- tdt l@&ori Uvarov,ldcn. rbcd., 2Al t63. 

t931. Ptnu@bius corcolor Choprd, Butl. RrfflB M$., 6: 134. 

1935. Pt @r,, us ru@i Uv.d, E6, ll: 8t. 

ttA6. Ptdna'bbia ldazln w. ,tcw Wll,,,tu,ZLn. 

h io0axy 6rfonn! !o ihe publilh.d dB6ipdon (Choplrd, 1969; Sr..d, 1989) of 

MeEllmds(m):5d 59 

Mrt 

3,13 0.3r 

!,83 0.21 

lsbmabad I 9. lux-gri Hrnnr rilc (Qucn ) 2 

4 !, 9JX-93i Matnn Cqr 4 d 2 9, IGIX-93; Zid.l 

t,Gl_3 

2,5-!_6 

3,t1.2 

,.ru.o 

t.r2.2 

3,U 0.11 

d 2 9, ltx-96r t^bcla I d 

I d, l0-lx.93r Bhagd (Toba

-l9l- 

Tek Siigh) I d, l9-vUl-94; I 9, 2l-VUl-94i ChdSa Mang! | d, 23-VIII-94: I 9. 

22-vlll-94i Frisalabrd I d, l2,vll-EEr CojB (Toba Tek Sinsh) I 9. zElX 88i 2 6 2 

9,29rX-E& H.{d hlm {Yehdi) 2 d | 9,2t-Vut-qri Had Ma6tr (si.I@t I d 2 9, 

u-vl'93: Mianwali I d, +X 88; Muzfiar$rh I ?,2Gvll-EE; M!@ I 9, l3-Vlt-93i 

Pakpanao I d, 9-VII-88; 1 d, ll-Vll"8Ei I d, l2-X-88i I I, lGVll-EEi 2 I, ll-X-88: 

RawalpiMi I 9, 5-Vlll-88i I 9, E-VlU"88r 1 I, qVIII,88r I 9, l4-Vlll-88; I d I 9, 

ls-Vlll-EEi I 9, l6vlll 88r I 9. l?-Vlll-6Ei I 9,2o-VU-EE: I 9,22-VIII-E8r I d, 

unlat lled, talcn frcn thc srudc s' collErion, D.Panftfl of Agriculru6l EntonoloSy, 

UnireBiry of AgricultuF, Fai5alab.d, Ptlisi.n. 

h is a vcry cohhod ed g€ually hydrQhilou spei$ found or snall Egeblion 

abog &. banks of rive^, strotu, wai.r chanmh, uDdd dqd lev.s and on ofcn swi6 

or $c wat r surfa@ tu sne tin . Ir i! alio a C@d Rid ed thus ldgcly olleled oi 

lights. 

AII rhc abow l@litis. Earlic.. s@d ( 1969) @rl@t d it frch Faielebad. Goja, 

Mianwali, Muaffaisdh, Pakpalt.n and Rrwalpindi. 

Choped (1969) nE ioEd rhar rhc nicrcpErcB rdB or rhi3 speies w@ 

slft, b( Oe Mt.rial from this egion sho*s l|ift r}|, co$tituc n@ rhd 50* of rhc 

totrl olbcrcd populatun of this sFcics. ft ir lirdnq add.d rhrr rh. lpcim.B otleEd 

rrcn mountaiB de 6rch dark r in ecmr.l cololEtion rhan $* jn pldB. 

CinE Drr.rorilr Vicla.J 

l9?3- Dianehobis vickery, cirld. Enr., 105(3):419424. 

Type. Encopt tu foipd wztka 

Th. @llcicd mat rial .niicly @nform lo $e publish.d d.scripion (vick rr. 

Io?l) ofthB g.nus and a,epresencd by 2 sp6B In lhu re!'on.

-tn- 

Key to the sFcts of Dihôbiw Yl&ery 

| . Antdior and media. f.nor. paE wnidrh, provid€d wi$ vo t .svcM blaclish 

batds. Mdillaly pdpi with ldt teo sshcnE nor whnbh. P6E id ribi& with 

3 exEtnal and 4 i lrnal spin* in feMls. ........... crrhn Ooh.r) 

_. Ant .ior lnd m.dian fem@ whnish in bul hall brewnbh in !pi6l half and 

wibour dy bul- M{iU&y palpi wilh l$1 two Fgm s whitir[- PosEriot ribiac 

wid 3 spitur o. both sid.s in fcnales. . - . . . . . . . . . . /@.4,.r (walker) 

r. Ditr.nobius .tili Oonv$) 

(Figs. 52A-D) 

l9ol. Nwbi6 .riki 8otiw, zichy, Zool. Ergcbn.. 2: 243, 

1925. Pte@.tubiB 6 rt Chopdd, Am. Mag. nar. Hbt., (9)15:50?. 

t9at. Dian.Mbi8 .ritn Corckhov, Vesm. Z@1. 0(2)r 2l-26. 

Thc @ll@t d sp@im.N loklly led *ith th! publilh.d dsigtion (Chopa.d, 

1925, 1969r S..rd. 1989) of thb spei4, with the addition of de followiDg ch$&t s: 

Th€ neropt rous femalB show tully devclo?ed clyFa, which dc rcund.d 

apically (Fi8. 524), shilc tl'c btuht?r.ou ford hav! t.n clyE! dcdinS a lnrb 

beyond lh€ niddle or abdonen .'d Mry nu@r.d al ald (Fis. 528). 

M€sureNnrs(m).2d 39 

E,L, 

n.u.r., 01,' otu

Fts. 9. 

-193- 

c 

Dlaa.@Utt .tiL& A, drb. oftr. &drrtqlB r r B, drh 

oft& b.drt.6r 9i C, paHa f@ ar D, !...t- dla a, 

B

lt.T.L 

-19t- 

0794.91 0,35 0.06 0,76{.92 o,34 o.o7 

0,63{.''1 0.67 0.04 0.5it{.63 0.62 0.06 

3.0{-r.ll 3.r9 0.r5 r_77t.N 2.27 0.16 

H.T.[,m.f.L 0.76i.79 0,73 o.O Oj'4JA 016 o.g' 

I|.F.L./O.L. l.2Gl.rta 1.32 o.to 

Shae Goba T.k Singh) 2 d 2 9, ICV I-94i Kh.eval I p, IE-VII_EE. 

It ha! b.c. @lt@0cd frcm the pt@s klr rotting logr, while I single 

o.dopi.rous f.hsle wa fou.d on lieh. Ed a culrivakd fEtd. 

llris sp@is hd ban @I@ted fo. lhe tiBt ri6e non Bhagt. pr.viosly n wd 

@lleccd frcm Kharcwal by secd (1989). 

2. Dian.@bits f6.*B (w*ai) 

1E69. Ereopt.tu JMip6 v/etk6, Car. De.@pl. Salur. Brir, MB., l: 6?. 

1871. Eneoplerc altoam (f. b@htpt.b) walk r. rbid., Suppt.,5: ll. 

1871. NtubtB ndao S.ucaure. Mem. Cen.ve,2i: e5 

1893. N.tubiB nietbientr8 B@nM, Am. Mul. c.ftE, J3: 196, 

1904. N.tubig Ditl@.rr6 Mr$unuE, Tlou. rs. Jap., tr l3s. 

1906. CJndipha atboana g. b@rt pteft) Kirby, Syno& Car. Orth., 2; El. 

t9M. c\nqipla lMipa Knby, rden. Ibid.,2: 81. 

1921. n rene@bi4 lucipet yin Chi Hsu, p.kin! Soc. Mt. Hi.r., BuL, 3: 30. 

1924. Pterch.Nbi6 /r: nb Clmpdd, R.c. tDd. Mur.,26: l&2, 

1925. Pt tuE@bid atboattu (1. ha.n F.re) ChA&d, AD. Md. et, Hisl., 

(9)15:5m. 

1929. Nenobi6 f6cip8 C,F.Wu, prc. na1, Hist. S@, Fuldcn Chrisr, Univ., p_ 25. 

19?3. Di@.6obi6 lsci\s Viclrery, CaMd, E ., lo5(3)r419_424.

_195_ 

Th. dllat d nar..ial gi.{dy r*nbl6 ih. publilh.d dBcriglion (ChoPlrd, 

1969i Tandon and Shisndia, l92r S.ed, l9E9), but diffes fon rhc origi.al 

d*riprioi (walk.r, lE69) of rhi6 specics in rhe fotlwing chdcr.n: 

Oeiput with fu pale liB (dcly ab&d: @iltary palpi win bsl t$ 

kgdcnls whnilhi poli.rior tibi& in nab wnh 3.xErMl, 4 inErul lpin s: 6rci black, 

wirh a yelbwnh riog Dd. b.!e, 

M€aoftheds (m): sd 59 

s,o. 

t.50-2.6t 2.06 0.33 

L$h.la I d I 9, 9-tX-91; Maka. Ctut 2 d 2 !, !GIX,93; Abbonabad 2 9. 

l4{X,9r: MaMhn I 9. l5{X 94: H6d Batroki (Ksur) 4 d, l5,vt-93r Cltansa Mrnga 

(Kdur) 19,22-Vlll-94j ld2 9,23-VIIt-94tt3lanabad 2 d I9. ll.tx_94; Rlwal

-l9G 

L*e (Rlsdpidi) I 9, lGVIt93; Mu@ 2 d 2 !, l3-Vlt-93r pa&ctsl I 9, 

25-VII-E8; Pawalpindi 19,3-Vlll-88;t d I9,4-vtttSEt I ?,5-v[t-88i r d39, 

lo-vlll-E8; 2 d I 9. l5-v -88: I 9, lcvllt,88: I d, 23-vttt-88: I 9, 4,rX 88i 

Sanela Hill (Sheikhupula) I d, 22 X-93i Sukhcki (Fabalabad) I 9, 26.V,94; 2 6 3 !, 

unlabeued, ratcn irm dlc stud.nn' coltcctioi, D.pa.llMr of Agri@tiural Edomtogy, 

Unilereity of ASicujturc, Faislabad, Pakistan. 

It is gcfthUy hydrcphilols md coll@Ed froF snalt vcgeEtion along $e benkr 

or riv.E, la**, st@m, Mr.. ch2mh, und* d.ad lavB in foBtr ald fton rhc righ6 

x 's a.oomor 3paja, which wa @ll@d for lhe nrsl d@ from aU th€ abov. 

l@liti4. Earli.r, S...d (19E9) r@d.d n nod Raealpindi and pdqand, whilc 

Abdnllah (1955) from an uirnopn locality- 

h M 8.rerally obw.d during rhc pr* studies tur rhe sicrogt 106 

sp@inca of this sFciB werc more @nnoq will rh. f€oatc etyra alodr Elrer. 

.p'crlly s @mpared wirl' Oc rEcroprc@u fchalB, eirh their €lyra rcdly routd.d !r 

apex. Fuder, U€ speineB coll@lld frcm high alfiuds e slch ddlcr in rhcir 

gcneral bod, @lounlion rhan rIN Aom plaiB. 

Femily Trigonidiidle 

6lal nunber of four sFciB dd subsFciB hs b.€n idcDrified undq th@ 

Key to tlp gum of ftigoridinb. 

l Elytra glabrcu or alnosr elabroB, widi veinr alhGt p&alcl (Fi8!. j4B, 55) 

Elyta cole.ed wi$ firc 

(Figs. 53Bc). panicuLtly 

pubes@Dce, wi1h veiB Dorc

-t97, 

2. Body bl&k, with Lgs yellowirhi cly@ 6rnerus, ulh 

principal v.iB (Fi8. 548). 

Body and leSr palc broNnish-ydlow; 

bet$eo lhe prircipal veiN (Fig. 55)- 

dyh mcmblfuG, *irhout frle v.iN 

cffi Awtut B'l)|'@ votr wrtldrtl 

1E93. ,liutut|A Btunftr v- w., Ann. M!s. mt. G.mE,33:212. 

1925. Ponnariphd Chopard, Ann. Mag. Mt- Hi!r., (9)15:530. 

T'!p.: Ahuuteut ttlvB Btnnnq \-w. 

Chopdd (1966, t969) @Bidc..d A^*48 !"tea BrurM a tt symnym 

of E4eopt.re oc.!%ioa.t walka and rl$ dai8dt d it 6 the 0?e of th. 8.nus. 

Bu1, co.fgry b it, O1t. and Aldindci (1983) ncntioncd that e udMid8 h hoo 

Ceylon rd .,i- Jtlvd frem srh.rn Bom, e rh.y @ tut bc thc ry@!yn! de io 

tn€ disbne.nd wter bdric^ involvcd b.rw@n thc two t,?e l@liti6. 

Thc @ll@Gn spein B ofa siD8l. rew sp@id evcdy @nform to llle publhhcd 

dsciptiotr (Chopdd. 1969) of this g.iu. 

MALE: 

1. Aûsurgtt kmtu6 sp, N, 

(Fiss. 2c. 534_D 

SDc small (5,75 - 6,5 nn). Body @loution ra$er unifornly yellNhh. 

Hc.d 0,8E - l.l0 tiro d widc a nuinun {idth of prorctls &rG .y6. a 

litd€ slopinS at venex, previded with r.ddish poinB, @h b€ain8 a brisd.i ftontal 

osrum n4rowly prcdu4d berwen tic ant€me, shoner than lst a ennaljointi f@ 

and chcks cflircly ycllowit. Ey6 palc brown. !€niolly lensthc!.d, st olgly bulgin8. 

Artenr trd p6lpi ycllow,5thjoi of dE @iU.ry longerthd4th, disriE{y eid.ning 

ar ap.x, almo$ iridSular. 

Pronorum 0.55 - 0.70 tin* a loog d vidc, feebly drowing in font, il5

-t9& 

anterior margin straight, posterior produced brckwards in middlei dist sligbrly convexi 

unifo.mly ,clloeish, provid.d wi0t fin. pub€ge. qi|h small rcddisn tub..cl4, ach 

h€dng a brbdc, panicululy alons pct rio. mdgin (Fig. 53A); htlnl lob6 of rh. s.ft 

Le8s 6iobou! wilh $e body, dcNely pub.sntr ant rior tibiac slighdy dilated 

at be to a@odod.c large, oval rymp.nun on @h sid.: pGrerior f.sof pale 

yellow, elon8at€. MrNing towrds ap.x; posr.rio. ribie sl.ndcr, poid€d wirh 3 

deliare. pub*ent spirEs on ach side (Fig.2C)t i tetual apic.l rpus long, exrcrnal 

very smau. thc me

.19B- 

4.?4.35 4.?6 0.05 

s?.9.0 s.9o 0,r2 

r.2-r.35 1.25 0.06 

0.414.65 0.5r 

4.t5.0 4.77 0.21 

r,0,1,15 1.0, 0.06 

4.3-5.2 5m 0.16 

0.39r.r0 r.o3 0.r0 

0,5t-0.70 0.64 0.06 

4.915.39 5.16 0.lg 

t.92:r.67 2.26 0,31 

l0r-r o r.05 0.01 

Ilolotrp€: I d, t5-Vt-1953, Dapttr. 

Allotipe t 9, t5,Vt1953, Daphar. 

Pdtrt$ 3 d, l5-VI,1953, Daphd. 

All lbe specihea have ban htcb from rhe sludenB' @ll4tioni D€p&tm.[ of 

Agricukual Enbnology, UnivcBity of Agriculru.c, Fairtabad, pakisria, 

Disli.guisbi.g drrerc: 

h h$ a close reeemblanc€ 6 A. ozd@roid6, bur lh. geniBlia &e quit 

differ.nr and close to d@ of ,,1, /rd.u l!r. fton wiich i! en @ily be s.rarar.d by d. 

following .onbiMrion of chder^: 

F@ and chcek unifornly yellow; pionotun with tai.,al tob6 con@loroB wirh 

rhe dhk: elyra vnhour laErd d&k b!dr; gcnihlia (FiCs. 53ED. 

The sp.cim.B of rhir st@ks w.rc ptuiovty iilfJarirqds A@ipha brtipzans 

aiA. nlonotat by At/dutrstl (1955) aid sa.cd (t9s9) r..periv.lr. But rhc prene 

of finc prbesc€@ on elyra, absene of ctaccut nirrcr and anrl field, morc or less 

andtohos.d vcMtion of$e doetttctdand maL g.rii.lir slDws thlr &ru:lty itc

-2W 

F'a. Al. Arassttg Mat * N. A, [.d rd 9.ute 

d, tlo.il 

ac.d; B, dnn Jr c,lh. .D 9i D, o'ltdr.rt E, F, Dhr. 

@el4 aodl rd Err.l I'.dr.

:20t 

rpEimeE qcrc ofa Dw lpccis ud.r lhe Senus ,in&r!€6. 

Tl s spaies hd ber nm€d drcr ia body @lourarion, which h onifornly 

The type mtef ia! of thb neq lpcci* hd ben deposio.d in dr IMct MGUm, 

DQdtme.t of Agricuhnl Ento@logy. Univeaity of Ag.iculiurc, Faisdabad, Palt6l, 

G€nc lrtr@inu, Rlnbr 

1839. TnSohuiam R@bur, Faue Anld., 2: 39. 

1839. /t@b S.dille, lis. Onl-, p. 350. 

1853, Ce.atinopt.M Ficber, tltos. 3: 232. 

'lr"p.t Tngonidi6 ci.ind.loi.l6 R^nbrr 

The @llet d oal.rial ftom $i! rcgion conprising i sp.cks and a subspeiB 

ex&tly resemblcs the publbhed ds.iption (Cnopdd, 1969; Sac.d 1989) of rhb genB. 

K.y to lt sFci.. ol Tigoni.liun *Mb6r 

t. A erior tibiie unpdfoErdi clyra srongly @nvcx in la@ral pronb 

(Fig.54A). _ . . . . . . _ . . . . . . . . . . . . . . . . . . .a0d.ronh Rdbu 

_. Anr.rior ribiar F!f@r.d or borh si&sj clyrn dMt $nidr d slidrlr @nEx 

in ratral prof:lc (Fig. 54C). . . , . cirtrdelou.t ,lnrr'diuuD (SrNru) 

l. Tt goni.lian .i.itul.bda RMnv 

(Figr. 54ADD 

1839. TiSondit n cici4d.bid.s Bnbur, FNac Annil-,z:39- 

t839. Tnsondi@ pa&dlcola Seryillc, 16. Oni., p. 351. 

1853, CerotinopErus .icind.loidet Fi.be\ lar'os.3t 232, 

tE6[. TnconiAW o/@pfd@ Slal, Eugeni.: Rs, Onn., p- 316. 

1a6o. Tnqoni.litln nltrirL Shl, Id.m, Ibid-, p, 316. 

1469, kle/opt.tB .r.. Walker, Car. Dertupl, Salhr, B.ir, Mu., l: ?4, 

rE1E, lttu aipha ci.i"d.loi.1es guia.@. Sasurc, ldcn. Ibid,,25: 4?3.

ftg. 54. 

D 

.i.iutalouai A, afir a' Latn 

olrr.r &nd ..d Etr.l 4.dt, 

B, drtn 6, don l tlddt c, tL dt' 

B

-2Q3- 

1418. Htuediphg cicind.loid8 hanbenian|n SaNslre, ldcd. lbid,, 25r 458, 

ln8. Hotucuipha .al@pr'u&a Sausurc. Mem. S@, cercv., 25:472. 

lE1a. lb@@iph6 tlrtit Sausu€, ldem. lbid., 25: 413. 

l8?E. fngonidiun z6dedr@ Sarssur€, Iden. lbid.,25:464. 

189J. Pkttuiphw ririolB K&sh, B€rl. enr. ZB.hr.,38: 152, 

lE]6. M.tioeh. ci.itldcloud guin Mc Kirby, Syrcn. C!r, o.th,,z: 78. 

lE)6. M.tioehe cicindtbids hanbaia'un Kirby, lden. lbid.,2: ?9. 

lm6. Metioehe coleoptrutun Kirb!. ld.n. lbid.,2: 79. 

1906. M.tioch. riairL Kirby, l&8, Ibid..2: 78, 

lv)6. Maio.h. dr'ial, Knby, Id.n. lbid.,2: ?E. 

l9t)6. Trigori.titn /rttum Knny, ucn, tbid.,2t 11. 

1925. Ttigonidim cicindcloida &ir..6e choPdd, Bull. Mus. Hi$. M. Pd6. 

p.445. 

t92s. Tngnidith .ici Lbid.t hattb.ni@M ChoPard, Atrn. Mag. nar' Hist.. 

(9)15:525. 

l9y. PiBldiphs anbL. Uvad, Lini. S@ t. zool ,36: 594 

The colle&d spccincB 8@tly agrd wi$ lhc PublislEd d*ipdon {Chopad, 

1969: TaMor and Shishodia, 19?2! S@d. 1989) of this speics .xcepl the followins 

ANlrior and ncdian feson yellowish, sith a Eddish tingc in .pk!l harri 

posterior bbirarsi ligbl oloure

-204- 

1.7-2.0 

Mak.rn C@5t 2 d 3 !, !FlX-93i tsbela t d 3 9, 9lx-93i Panjsur I d I 9. 

.}lx-93: Abbo[abad 2 d I 9. l5-lx-94: P6hase I d. l-vlll-93i 4 9, 2-vlll-93: 

TarMb (Pcshawu) E d 6 9, l?-lx-94i Vek (P6hawu) | 9, l-Vlll-93i Cfianga 

Mansa (KN!ur) I 9. l5'vl 93i I d 3 9, 23-vlll-94i Faisalab.d I d I ?, l2_vl-88i 

Rawalpindi l?,7-vlll88rldl9, I l -lx-94r Sargodha 7 d 2 9,6-x-8EiBadin I d 

I 9.24-X-93i Karachi 2 d,26-X 93i Sukkur Bairage I d I 9,20-X-93. 

Thc apeimec of diis spei6 have ben @lleEd from s@ll gtds6, l@ itl€i, 

Egeabl6 rcd moirt pla6 ard lights n@ cultivar€d al6. 

h E vcry @mnon a.d wid.ly distribucd sP€ci6, which i5 rccordcd ror the ns! 

rimc from all the atrove l@aliti6, Earli.r, S@d (1989) @lletcd I ftom Faisalabad, 

Rawalpindi ard Sdgodha, 

2. I:tico'.Aiun cicw.toas hunbetlid'n ls.Baftl 

(Fi8s. 54BC) 

r81E. Hoha.onph8 cicir.l.loi.l6 hanb.niMun sa6surc, M.n. sc. G.n ve, 

75.468. 

\9n6. M.ti@h. cici".l.loid4 hunb.nian@ Kirby, Syrcn. Cal Orti ,2r ?9 

1925, Tripnkttun ciciadcbld$ hunb.niu@ ChoPard, Ann. Mag. n.t Hist. 

(9)15:525. 

lr largely r.Fdbl6 ttE plblish.d deriptoi (ChoPUd, 1969i Saad. 1989) of 

T. huhb.ni@un, erc.p. tl,e w ings which @ ontunl, .udin.ntary.

M*u|m.n6(nm).5d 59 

-205- 

M.*m C@t I d I 9, lelx,93i Ahbotl!!8l I 9. l5-X-94; P6hlwr 2 d, 2- 

VIll-93;3 d I I, l6'lx-94r Tamb (P6hawd) 2 d 3 9, l?lx-gar VaBar Osluwu) 

I 9, l-vlll-93; ChanBa Manga {Ksur) I I, l5-VI-93t I d, 2l-vllt-93i cojia (Tob. 

Tel Singh) I ?. 9-IX-EE; I d, 29lX-88; He.d Mlrala (Si.lkot I d. l?-vt-93r 

Palpaurn I 9. l6-VIl 88i Su*ur Be6ge t d t 9. 20 X-93. 

H$t.t. 

It is a conmn !p@i6, vhich i! msinly collccEd Aob r|ig, wg.l$l*, l.d- 

littcr nw noilt plac6 and fiod lights near te cultivahd fFlds, 

It is wi.hly disEibur! a'd B ollerd for $e 6Rr lin tion aU thc abov. 

lcalni6. E!li.r, saeed (1989) rc@dcd n frcn Coja and Pakplttan- 

Chopdd ( l%9) (thl! fauE % wrirEn much bcioE 195€, blr publLh.d in 1969) 

and Sed 0989) raEd it a a sepdalc rFci.r, bur 6e fomc. in 1968 lowercd i$

-206 

slalu to a 3ubspei.s of r. cicidedd.s. dn to its ct@ @nhtre in a .unbs of 

chardr.E including rhe geniralia lo rh. talter sp.cies, 

coN drc4,,t SrNun 

1814, /lnaipha Saussure, Mis. Mex., Onh., p, 3?0. 

1818- ,1|6iph6 SaNurc, M€n. Se. Cercvc, 25:475. 

l9o7 . CJ'lqipho Kmy, SB. At. Wiss, Wi.n I l5/l , p. 264, 

T]IE: A.hera dleao S^y 

A singlc @ll.crd f.malc ei&tly i'lli4 wirh dE publbh.d dedption (Choped, 

l%9: Ott and Alcxaider, l9E3)of$issenus. 

l, Ar6ipha loneiN^nb lsalllal 

(Fig. 55) 

t839. ThSonidirh tar8,iNM. Scdill.. lB. Oni.. p. l:.. 

t818. Ctndiphrs lrrsA?rrir Sau$!e, Mcm. sc. Ccnev,25:464. 

t$E. qndiphG pdr'lar Sr6ur., Idcn. Ibid.,25:486. 

1889. qdxiphut p6ri(a Bolivd, An. Se. .rpan. Hi!t. nar., t8:475. 

1893. qndiphus rrl4drd6 Brun..r, Ann. Mus. Oenova, 13:211. 

t906. Cyndiph. l,,a4r.e6 Kirby, Syrcn. Cal. Orrh.. 2: 8l- 

1906. Ctrtuipho r'arilla Ktby, Iden, Ibid., 2: 81, 

l906. CtnEipha ttr@iarb Knby. td.n. lbid.,2: 82. 

1923. Awipha pBilla B!(on & Uvaov, 8ull, Sa, R. .nr. Eg,, p. l8l, 

1924. Atl6ip\a lou8'r?rrir Cbopad, Ra- hd, Mus.,25: 188. 

A singl. ollaEd rpeitun onpl€lcly conforE io $e plbtish.d d*riprioD 

(chopard, 1969i Otc and Alcxander, 1963) of rhis sp@ics cxcepr in m6urcmcn|3 ol 

M.du@ots (m): r 9 

[.L. II.w, P.L, p_w. E.L. H,F,t, F,r,W. tr.T.L. O,L, 

t.0 a.0 

ll.wtr?.w, P.LJP.W, E.L./P.L. H.T.LJH,eL. II,F,L./O.L. 

0.67 3,39 Lm 2.35

-2gt- 

Flg. 55. A4siph, lortlp.^ntt, artn e.

-208- 

(BalEw.lnagar) I I, l,t-VI-91, 

from wild 8rd6 grcwn n@ dmp plag, 

Tnis sp€B [a b..n r@rd.

MALEI 

-?09- 

l. t|IoboAhd Mutonot4 sp. noj. 

(Figs. 21,56A-F) 

Si& latg.04.6 - 18.5 mm)_ Body otoudtion dek brNn. 

Small, d.rk b.owni @iput vith 3 brcad, ycltNish baods, wirh t*o anErior 

yellow'sh 3pots on dorsum: verFx stopinS, povidcd wnh 3halt bt&ti$ b.jstta up b 

iiontal rostrunt the latrer much udover rtDn lsr anlenMt segncnr; chccb and fae 

dark shinirg browtr, 3morhr dyFcfrend sutuc febty @ndcd. Ey6 jnongly 

prolruding, vali€lly leng0encd. btact, Palpidrk broM, thc naiill&y v.ry lorg, 5rh 

*8n.nr widcning ard obtiqocty rlncatc at ap.x. Anrcinae wirh first n! egnenB 

bl&*hh, whnish up b lhc nitdtc, rat brwn. 

Promrun brMbh-black, pubcsm, irs anrcrior half prevlt€d wirh 4 targ. 

bownnh-l.llow lpor llong the Fedian turiow, p6Lrio. half wit 5 ,m.ll. rounded, 

prohineni3pots in $e forn ofa cuRc (Fis.56,{). anrerio. maryin vcry sti8hrry 6n@v., 

p6rerior alnsl sraighr, bolh pdid.d wirt sda brbrlBi t.rdt tob€. of th. h. 

colou.6 ihe disk, wirh angl€s athosr rcundcd. 

Llgs long, sl€nd.r, of rhe sn 6tour s rh. body, all feDm. tilrt€ liShEr 

towards b4s; anterior ribi@ brown, prclidcd with sma[, ov.l tyopanuh on both f@: 

frrddleaM poeetior dbi& patd ye owhh, wi$ bes and api* browrtpGcrior libiac 

dm.d wirh 3 rm.tt, pub.$ent spi6 otr borh sijs, inl.rnati . linL ihdid Od 

externals. irrcrnal api@l lpu4 long€r $an exrrmls, medioinrerMts 0,4 rihes s tong 

s pos0ior bairnRus, th. tan r etongar., fe.bty @mpNcd; E sat ctas mr 

ElytB ext nding much bcyond lhc abdomen, vcry broad, 0,58 rinB 6 b.oad d 

long. dek brewn, wirh tighcr veiN; minor reIjev€toped, 0.71 - 0.?8 tiB s tong 

a wide, divided by 2 , 3 veiB, de hrter srrc,gty converging rowtrds ext rDl anBle 

(Fig. 564)i lsr chord angutai. in lhe niddt.. lnn€d wirh rh. dino, by a smau !.in,t

-21{L 

anteriot ar8le. 2ft1 rcguidly hnded, 3rd aln6t 3Eaighti halpi *irh 6 - ? oblique 

veim, nBt 3 'dly p&allcl, cunrd ar bsc, oth.^ sinuEr api6t ficld sho! pa.nting 

corfused vcBtiotr, ells ir.gulr; talehl fietd targe! panly @erin8 rhc body, Sc wirn 

l? - 22 bErchd. Wing. shorFr rhan.tyi-a. 

Dark b@wtr. C.rci y.llry, pubcsir, 0.88 rin6 $ lolg s posrerior fcnm. 

Cenitalir wnh epiphallB rilob{€ at po crior margin,lar.hl tobs long, robusr, mcdian 

shon, blundy poincd, sliBhdy cuNcd ar atex; ciopmm.r* n.dificd i. rhc iorn of 

loiS poin€d hooks (F gs. 56DED. 

FEMALE; 

It difi.6 fron $c oal. in lh. follNing charu86: 

Prom$m wnh posErior 6aBin febly conv€x. Elyt4 blactish, with nied 

yellowish veiB, Cu dividcd tE Diddlc. r.asvcrsc veinlds tru@rs, yellmilh. moc 

or l€ss.nastonq.d. fornirg irGella cclls (Fis.56C); latcalfield wirh Sc bdins lt - 

12 v.im. Wings long, 1.67 rima .! lorg a clyra. Subg.niral pbi. ft€bty mrtEd. 

Ovip6itor long, strdght, sl.nder, with apici] vdB mrow, snooo, l@lare aid 

Mes!ren€dte (m): 5 d l9 

B.L. 

3.&3 

,no

-2l|- 

Eolotper I d, l5lx 94, Abbona!.d. 

Allotype: I I, l5-Vlll,88, Ramlpindi. 

PlntJDs ? d, l5lx-94. Abb.ttabad, 

TIE mal.s of rni! sp@i6 *crc collctcd .t nieht frcn a grury lawn whil. 

suidllating and a singl€ fcn le from ligh nqr a deae for6t pLntalion. 

DililtrguiititB ct.nd.6: 

It hA a cl* rescnbl@ to M. japo ca, from which it can 6ily bc 

separgt d by 0'! hsd aod poootal Mking!, anien@ brown in dktal half, mkror 

divided with 2 - 3 reiN, Sc bdins 17 - 22 barchB and stiihlia (Figs.56DED. which 

are closer to thol. of M, ciacti@mit, 

Tte arlorype fcn L of thn spei.s B idcntificd and .cdcqibcd I H. jd?oricts 

by S&ed (1989), but, id vicw or the above differcnces, il appcarcd 10 bc a ftw spcciB, 

wtich is sinil[ io ma16 @ll-ed from Abbou.b.d. 

The .ame of thb spc.ies ha b4n ddived fron it pronotun, which is providcd 

with r baulitul parFrn of y.llNish !pot!. 

The type roEriel of rt! sD.!i.s ha b.!n deposiicd in 6c Insdr M!*!m, 

D.pann.nr ol Agi@hinl Enbmoloty, UnircBny of Ag.tllruE, Fatalabad, Paki$an.

Ft. 56. 

z c>.-s \ 

#r= 

o7o, 

M.tolturpha M.ulo@la .D. e, A, haddd Dr.!a@ d, 

do.d.rd; r,drrr,; C,lt s 9; Dr, aEt'r. d, CLttt 

o'e|d'illnr|'Ydrd.ndb{.rl!'.d!.

FEMAH: 

-2t3' 

Z. M.LtnoQhd amlArz sp. nov. 

(Fig. 57) 

Sia m.dium (13-5 nn). Body olollaton blekish+rwn. 

Snatl, bNnt @iput lighr colound, wnh slighrly visibl. 3 teltqth b.ndsr 

ve ex sloping, mert with ltu trllosish spots.nd small brisll*: fronral rdrun v.ry 

lurcw, qith a row of bl&k b.htlcs on int rnal sidc of @h antenml sock€r. thus making 

I fumsedi ch@ks dark brown, sm@th; face broNn, wi1h y€llowish tiDge rt clypcBr 

clypecfontal Nture feebly rouid.d. Eycs black, distinctly prcrudinB, vedc.lly 

leoSdeied. Palpi da brcwh, nnely pub€s@nt, lhe @ill&y rcry long, witb 4th 

s.gnent slighny shoid rhan 3rd and srh, th. lattcr Prdeniry, and obliqwly trlmated 

at apex. A eo@ vili fiRl t$! joiil3 bl&kbh, rioged widr al@re brwnhh aM 

yellowhh ir rhe bel pan and apiql hdl Bt whirilh, 

Prorctun tap.dng dcidly, d&k brcwn, Pub.snt, 

iE ui.rior hdf eilh 4 

yellow sp* along nedian liH, Dosi.rioi halfwith 5 snull. reunded, obtuF spots in th. 

forn of a cune. aturior marlii co@ve, porloiot fcebly onv€x, bolh pr@id.d snh 

a row of snall brhdsi lateal llcld detp broqn, ending posEriodyt4s 

sligblly lighter than body, pub*ent, aU fenolt ddher in apical lEli 

antclior ibie brow., Pefotatd 

with sBall, ovaltymplnum o. both sid6; niddlctibiae 

ycllow, with bases and aPic* bown; Poterior fcmo.a sl.nder, wnh filifotn aPidl Pel, 

ribie dark brown ar bde, ..d in apical hali .csl y€llo*hh'brown, emed wnh 3 smll 

spin.s on @h n&gin, lst itt rMlv.ry3mall: apiel spuB 6, cxLrml I short, inl.rul 

3 long. mediointerMl spurs longe$ on 4h side, 0.4? rins d long d p6!.rid 

b6iu6N. rhc latrer onh rupqio| rur8in d.nticulet d. 

Elytri @Ner, exta.diig kll beyond thc cd of abdomn, bl4ki5n. wnn 6i*d 

ftlowith vciN, nsv* rcinleE Is aBomos.d forning 

i.rcgulu etls in batal lElf aF1 alhost r.gulu. clongat @Us i. aPiol half (Fig. 57):

:r,l+ 

It, 3' ,W e t '.arne.

-215- 

latenl field virh Sc having 9 - l0 bhrchd. WinSs lotrg, r@ning rhe rip of oviposiro.. 

0.74 tirne! a5 lo.g as the elytr. 

Da.k brown- C.r.i y€llow, pubescent, 0.94 tiN a loog d pdlerior femom. 

Oviposnor lo.g, nraighl, with rpical valvcs sm@th, Mrcw, ldeolaoe, @ut ar rFx. 

Mesurem€d(! (m): I I 

P.W, E,L, W.L. H.r.L. H.f.W. ILT,L, O.L. 

2.5 2.0 3.75 13,5 20,2 1225 2 r5 t2.5 

lt.w./P.w. P.L./P,W. E,L,/P,L, ll.T.L./Ir.F,L, x.F,LJO,t, 

0.67 0.53 6.75 t.Ol 

[olotne: I l, ll-lx-94, hlambad. 

A single female s @ll.cted f.on light D@ a deR plo|rtion along .@d ride. 

Distitrsrisnhs chdd€s: 

ft is very close b ,{r. MtloNtd,sp. Mr.lrffi*hich it en only be spmEd 

by rhe a.cnMe, which arc disti.ctly annllaLd vnh yellNbh a.d br@h, oboe 

pronoul spoE and slighdy sn ller sie. 

Ir also sfiows it afUnity to M, circticodis a.t M, baSi@ait, Wr dittec 

nom boih in hed and Fonotal markine!, wins length and nea$re@na ofveioN body 

Tn€ Dame ot $is spe.i.s hd bccn d*ived ircm ia a €n@, which arc ainulated 

with brownish and yellowish. 

Th€ hololrT€ f€oale of thb spei.s ha b@n d.po3itcd in $e IBet M!*um, 

Department of Agriclltural Entonology, U nivcBny of As.iculrur., Flbrlabad, Patbni.

-2t6- 

tr'amilJ En€opieridoe 

lr is repr.s. .d by rtu sp..i* undcr wo gemr. ad two subhmitics. 

Key to it!. sublilniti6 of EneoDtqtdre 

l. Poslfior ribiac wirh mod.rar€ty tong dr.rmt.pi:r sius, rh. mcdian much 

long.r thln orh.a (Fig. 588); pct rior bsitani tong, M.ty d tong d ,! rcxt 

rwo *gochls rogerhcr. . . . . . Eneopte.ioe 

_. Posterior tibi& {irh !hon, subequat external apical lpuB (Fig. 598)i p6r.rior 

banasi grarty shorcr rhatr de rext rm s.gnlcnts logerh.r,. . pod06.irritr& 

Subfsmily Eneopterln!€ 

The cou€ct d spaiincns fau in a sin8te spccks uMd a sinSte g.nB. 

c.nus .Xrrr8rr&r Botrs 

1890- X.nogryUu Bolivar, Jorr, Sci. Lisb@, (2)l:232 

1E93, DionlM Brunftr v. W., Ann. Mus. Cenova,33: 213. 

'fypct Xeno|ryuw .r.opt.tui.la BFitiyt 

The @llcrd mrial of a singtc spdis trcltly .6embles $e pubtished 

d*riprion (Cho?atd, l%9r Sa!.d, 1989) or this senu qc.pr uE nKori which is 

divid.d by an anlulac v€in (Fig. 58A). 

t. x.aosry a, ranr.d6 (w.lla) 

(Figs. 2BF, 58A_C) 

1869. PlatrdutytB rrdrrv.frk watker, Car. De.mpr. Satlal. Arit. Mus., l: 88, 

la7'. Cadiod@ryLB ,r@yr66 Sausurc, Mem. Soc. c.ev, 25:525. 

1893. Dio.rM .aLrad Brunn r v, W.. Atrn_ MB. c.neve,33r 213. 

1906. MadNtmna rr.tuvdrub Kirby. Synor Cir. Orrh,,21 94, 

t%8- xe@tryt\s 'arrar66 Chopard. Orrh. Cat.. p.rs. lO, cryll., p. 350. 

n @mtlcaly .gB widr $c pubtished dcsqjpri,on (Walkr, rE69; Chop.rd. 

1969i Sa€€d, 1989) ot dis spdi6 except rhe Sc, whbh I pbvided widr l? - 21 

b.amhcs in nates.Dd t2 - 13 in the f.mat*, Ceniulia (Fie, 58C).

t. 5r. 

:tl?- 

M.enmn.s(!M):5d s9 

-2ta- 

,r.oo.o ,;n o; 

hlm.bld 8 d | 9, l0 tx-94i Ci&ga Ma.gs Gsu) 3 d, I 9, 22lx_94i2 

23rx'94; Mirha Tawana (Khlsbab) I d r 9, l4-x-91: p:*ltpitdi I d, 2_vlt-88i 2 

4-Vlll88; 2 9, s-VIll-8Aj I d, 6-Vttt-6E: I d. -VIU-E6| t 9, tE-Vl[_ESr I 

l9-VIIl"88; I 9, 2l VUI{6i I ?,4-tX-8E. 

It v$ collered fmn densc bushs and sdkanda (L../tarunr ,/.) white sinsint 

at nigh. and on li8hrs n@ lore* ptai'rios. 

All thc above lqlilics erepl Rawalpindi, f@m whm ir ss prcviously @dcd 

by Saccd (1989). 

9,

-2tv 

Subfamily Podoocirtina€ 

lr is r.pr* rd by a ringlc spai6 uider a sinSle 8e.us. 

C.nc Ercrrtlr CHin-M.revlll. 

1844. EBqn6 Cu€rin-M.mvill., l@i. R. anim., lns.. p. 314. 

1814. E6cin6 Sau$uE, Mis. Mcxiqu., Onh,, p, @, 

'typ.: Euytdu riutldru Crcin-McEvillc 

The speiocN oi a si.Clc ap€cia @npt€t ly agre whh lhe pubtishcd d€scription 

(Chopaid, 1969) of thb geiB. 

l. 6at!,1us .oaciarus (HranJ 

(Figs. 59AB) 

tU2. C,rU6 (En.opt.ru) orcrrr$ Hdn, Tcmoirck Velhlndt. Oflh., p. 231. 

1874. EeinB .r.ciaB SaNurc, Mem. Soc, Cnev.,25t 626, 

1889. Eactnw ."d@6 Botivar, An. S@. span, Hist. qr., t8:430, 

It r*nbl€s wcll wilb rhe pubtilhcd d6diprion (Chopard. t969)ofrhirspei.s. 

but diftls in rh€ followinS chaact sl 

Head son ri@ wirh prominent brown bands; po3rcrior fe,noa pale, dillus.d 

wi$ browD on exr.rnrl fM; pct..ior ribj& &mcd wjrh 7 , 9 .rErMt, E - u inr.rnal 

spi.es, with inrcrEl mgi! fd rhe mr p|n having srnt*ioB b.rs@n ihc spiB 

(Fi8- 59A); clyh brownisn on m.diao ti* in d{tc. speiDcN, 

M.a@mnar(m).5d 5! 

B,L,

.* 

Il& t Wt .,/rtutt \@6t drE,.F.?bry, 

I

-221- 

Tahab (P$hawd)'t 619, t1-tx-g4t Vdrat (pcrhawd) I 9, l-Vflr_93l 

Chahea Maiga (Ksu) I d I 9,23-vI94; prwd t*e (RasllphdD 5 d 6 9, 

rGvrr-93:8 d r0 ?. ll,tx-%iAylb pdk (Rawalpindi)2 6 2 g,g-vlt_%i,t 619, 

l0-lX-9r1. 

It w.s foud in dleiation with giasshopp€s and @lle.Ed frcn tong grag 

.ror8 *arer couesi ponttli srens, .a_ 

Thir spei6 ha b€ln r.@rd€d id rhe tiBr rioc fron patBhr 

Famity O€crnthid!€ 

rr a ftp.*ncd by rhce sp€ia ordcr a singt. gcnB. 

cenls O..o.rntu Sqtile 

1831. Oea,tth,.t Sdi[€, Ann, Sc. iar,,22:38. 

1435. Aeaithw 8tu ., Hist, Nar, tN.,9: I74. 

1837. GFJllon ia Scidt, Weit nrveb.r B€itr, Ce!, Natut. H.Iwis,, l: 212, 

.fr,?c: Achaa ir.ti@ F&ricir, 

It. cott.cr.d iFcireB exaoly la|ly wirh rh! pubtishcd 

1967; On€ ard Al.xander, l9E3) of $t 8e,us, bur diffcr koo 

S@d (1989) ii th. nirrcr, which is divid.d by 2 veiB.

l. 

2, 

222- 

Key to lhe rp€cl6 of Oacdrrr8 SeFill€ 

(14.6 nm or ls)i sridltaro'y 6tc rerh 33 - 3?r t.ngft of ctyra 

lN. - . . . . . . . . _ _ . . . _ . . . . . . . . . . . . hdns SrrssE 

(15 Fn or norc): sridulatory nle t €tn 43 - 53i rcngth of elyna 

..................... 2 

Stidul.ttrt fil. te..h mE di.r 50: ,honcn wirhour d-k b@n mid{o6al 

stripq B.nir.lia with mdid lob€ rctchc! al apei (Fig.62^r. ...,..,... 

. . . . . , c.e|ottlus 6p. \or, 

l4s tian 45; abdomcn *ith d.rk brown mid-dorsrl slriE: 

lob. mr mahed (qln6i iu-aisht ar sFr (Fig. 64/{). . . 

........ d.ntotp6u tD. nor, 

l. O.cadtar hdtls StNUn 

(Figs. @A,D) 

la18, Oe.otth6 r'rdi.6 Saulsue, Mem. Soc. cencve, 25i 454. 

TIe coll.cted haleriat entncty agr4s with lhe pubtbbed dscripdoi (Chopard, 

1969r Tandon and Shishodia, r9zi Saed, 1989) or !hi! speis, wftn $e addition 

Eyes febly prorrudins. venielly tetrgrhcnali ndiltdy patpi stend.r, wi$ 5rh 

joinr.longat€, longer than 3rd and 4rhi p*Erior ribhe arm€d pith 3 - 4 internal.3 

.(.rnar sp'Gi sr.idutabry vein pnided fith 33 - 37 reth (Fig, @A): tdmt field 

with Sc luving l0 - 12 brarchgr erci pub6ent, slightty linucr gcriiati! 

(Fiss. 6]0BCD) uirh cpiphattus rritobcd, mcdiar lobe li$ty 6drc{r. 

M€surem.nts (@)t 5d r9 

2.2

:223 

l2.5Gr5.O 13.94 0.99 

6.50.6.30 

3.55-3 $ 1.71 0. rl 

7.0G7.30 7.14 0.10 

0,3t-r.0 0.97 0.06 

7.5-3.20 ?.90 030 

r.55-r.?3 1.62 0.06 

0.9Gr.0 0.93 o.nt 

0.3+1.02 0.0,t 

5.I.6.00 s.6l 0.3r 

r67.1,3t 

r.05-r.09 o.0l 

Abborhbad I d, l5-tx,94r Ayub palk (Rawatpiidi) 19, l0lx-94r coja (Toba 

Tck Singh) I d,8-IX-88; Mutlan I d, l8-tv-92;2 d, untabelt.d, l.Ln fiom rhc 

nudenK @Uerion. D.panmefi of Agriculruat Enbmology, UniE6iryof Agricutrue, 

Tlis spoci€s was gemhuy cotla0ed fton tong grasra and shrubs, bur nrcly on 

lighr Mr a dcnF for€lrpl.nlalion. 

lr h6 ben recordcd for thc fiBr tirc frcm Abbon bad, Rrwatpindiaxt Mutr.n. 

E3rlier, Sac.d {1969) co €crld it ody frcn Colla. 

The endE @tt.ct d masial of lhis genB h[i$ with rhc pubtGhcd d.srhrio. 

ofA td,.1,r. B a carctut cxamiErion of dE lr.irutdory fit. r.€th ad matc Beniulia 

sbow€d th.t ir ws &hdly . spei€s @npLr. N* lhe probten *s which oE betonS.d 

to o. indiM, b€@Be nenhq rhe ds.iption of file Eetn nor rhc diasram of Date 

genitaiia {d av&ilablc for tis sp6i.s, HNev€,, trle body m6 

glandula! projection of a group of sp€im.ns g@tty ag@d with t6c of O, rnd,c6, 

to *hich rhe aurhq hd dsigftd Oe eme Mn.. 

,:" 

r,3l 

1.33

h,o, 

'274' 

.,\. ,/i 

))Ttttt4l;xrn'-tt::!-zirtr.fia,.* 

I 

A 

-r' D 

at et tu a.*r!.aitx,*.; 

B c, ,E o4&a adl .!d n!.1 rt .la D' F ot rb 

c

MALE: 

225, 

2, Oeceth$ cwottlut sp, aor, 

(Figs. 2DH, 6lA1], 62A,c) 

Sie ldgc (15 - l? mn). Body @lq6tion yellwth-whi0., slighdy tingr! with 

Elongat4, whnish, diffusd wirh light b@n on n.dian liE; frenral Grln 

namw, slighrly Mowcr dEn l$ driml s.gmenq fa@ vhir[h; clyp.o-fiona] suturc 

eligltl, rounded. Eyes fe€bly proruding, reddish-brown, longer tha. wide, Mdillary 

palpi lons, sl€nd.r, eith tat sesmefl a hrtcswotlei |oMds aFr. AnE.ia. whirc. 

Pronotuo elongare. $ongly Mrowing in tront, ik daior mddn srlienr 

Pdt rior slighiy conv.x; disk alm6r fl . whitisht larlral tobcs smalt, iofcrioi nrSin 

sinuare, dcs@ndins pGreriorly. Meirnoral gt&dutar pit provided wirh r sFo.gty 

devel@ p6'.donydi@r.d angutd prejerio, ilc la .r @row.r ar bac, noch 

b.@dly rcunded at ap.x (Fis. 61,{)_ 

r4s slend€r, pate yeltowbhi anErior tibie stroigty ditated ar b.ee lo 

amEmod.te $. ertrcnely ldg., ovat ryopanum on borh !id6; pcr.rior f.noa tong, 

rrcnd.ri posler,or tibi& erEd wirh 4,5 inr{ul, 3 4 .xlc.Mt 3tuI spiB (Fig, 2D), 

aprel spurs 3 on @h side. the supdior longer on bolh sid6: epi@t jpurs. !pin6 dd 

denticles blact at apies; posreiior bsilarsi sroDgly conpr.s$dr tr6at ctaws 

Elyrra ratupocnt, p{te gBnish, qr.nding much berond abdomenr miftor nuch 

lon8er dan wid., routdd poEriorly. almsr angul& anteriorly, divid.d by rwo veiD 

(Fig. 6lB)i barps with 3 roatl veins; tateral fi€ld wiih Sc ll - 12 branchedi sfidula6.y 

lil€ proyid.d wirh 52 - 53 rcth (Fig.6lC). Wings tong, LIE - t,2O dnB s long 6 dc 

Tergit6 pal€ yclloqish i[ bsat hali, brcwn iowalds ap€r. Cercipale yclowBh, 

sfionEly pubegm, 0.66 tam6 d tong d porrcrid fenon, ceniDlia with medi.n lob. 

dis.rcrly mtched at ap.x (Figs_ 62ABc).

:zx- 

..1,, 

,1...,rainrr1iti;;rtii:ii9rti tli:iii+ rila4** 

c 

Fl.6r. M.t .!@tthtt t d. A' abatb td ., L 

trdra'.Fr ar C'-*t rd.

nl.6l. 

zn- 

m .trad!, + t'ra' af al /T'' fo@lr* 

&.rl d t-.t * C' F dO. E r.t 

I

M4Elqprs (|M): 2 d 

Eolottp.: I d, 

P.rrayFr | 6, 

-228. 

13.0.13.50 13.21 

l,tGl,59 t.55 

l5{X-94, Abbotibrd. 

Thc holotype oa thb !p€i.! w @tlcclld iion lorg gBlB in a v.gctable fi.ld, 

s.D, 

0,13 

shile thc p@9?. fron f.@ during lnidubrion ar nighr, 

Dblinru&llrg ch!r0.t6l 

It hd a d@ r.s.dblac b O. iidn6, bur ditrc$ filb n in blvilg rbe tdse 

b€dt siE, gr.ab Nnber of filc reth ard rbe mte 8.nitdb with mdiln loh. no!c!.d 

ar 8Fr (Fig3.62,\B).

-229- 

The na@ of ihis 3pei6 h3 bei dc.ircd lion rhe n.did lob. of iB e?iph.ll6. 

which is not heil at aper. 

Th€ type ma€.ial of dris sFcics ha! b*n depGitld in the l.s4r Mueum. 

D€pannent of Agricul$ral Entonologr, Univ$sity of AgricuhuE, Faislabld, Pakirlan. 

MALE: 

3. Oecuthut d*t tpa6ut sp, no!. 

(Figs, 634 C, 6444) 

Si& large (15,5 - 17.0 mm), Eody @lou. palc yellowish. 

Elongatc,slightly wider dEr anterior width of prorlorun, yellqishr ve ex wilh 

wo v€ry clos. loogitudinal rows of bben dors in th. middle, a sMtl blact lim behind 

DdErGint.rn l hargin of €ch eye; fronral Guun huch wowcr rhan Irt er.nml 

joini face paleycllowihr clyp@-frontal slNie rlnosr sFaiBht, Eycs black, ting.d wirh 

reddish-broM, venkally l€ngdEncd. Palpi pale, lh. darithry elonglle, slcnder. 

Anreo@ pale ycllowish rcd b6e. bro*n iowds apex. 

P.omtum.longac, wiih sr.igt ei.rior, febty coovq p6re.ion @gini dkk 

pale yellowish, sliShtly convex, with 4stil,like, f*bty vbible, ligbl reddbh-brcen sporr 

towrtdr l.t al cdgs rcd 4h @rEr; laE6l lob6 v€ry tow, int rior ourgin tEbty 

sinoate, M€ndiDg an|4riorly. Mclami.l glaiduts pit pmvidcd with . psr.dorly dn@cd 

l€ss strongly develop€d projerion, (hc laner febty bed.r apex (Fig. 63A). 

t!8s pal. yeuowiih, wirh brownirh ringe on po6r..ior femoB; aErio. ribi& 

strc|rgly dilat'rd n6 base, prolided wnb v.ry large oval lympanum oD both sides: 

potcno. ranoa v.ry @rcw, sl.nd.fi pscrio. tabi& {m.d wirh 4 - 6 inierul. I 4 

exrcrEl small spin6, apiel sp!6 6, slp.roinbnuls tongcr d.n orhcs; ponerior 

basiraEi srrcngly conpl*ed, so@rh,

-a3D 

. ,tr 

))r'*ruuiu'1-1tdt'*tI$I'fl 

lla3. l* E- +t-t*.d- 

-atl.i 

c 

ar C'.r15-4. 

ii\

:23t- 

Elytd nwh loryd thd.bdont ni t.nsporcnt, p.lci nircr much long6 0r$ 

wid., rouflLd pGr.riorly, divid.d by t*o Fr.lLl v.in ; drgpnal vrii srdgiq dodt 

with 3 obliqu? veim, thc last vtuy smau (Fig. 638); hi.nl fcld taBpent, of tha ffi 

colour a doBal field. Sc bdiig 10 - ll bBnch$; eriduhtory til. p@idcd eith 

43 - 44 r..rt (Fis. 63C). wirg! Err long, 1.20' 1.35 tiB a lotg |. tlE.lrm. 

Ycllo1lhh, fe.bly spotted with light bwn, dorun pwided w[h a ndrroo mid- 

doiel brewn bad. C.rci ydlN. pub@nt. loi8, 0.?9 tiB it loiS 6 po.Erior 

fcnffi. SubScnirl plar la4r, bwnilh-ydlw; gcnitrli! witb ncdi lot .ln6t 

nuMo at apet (Figs. 54ABC). 

Ms|tt|roas (@): Sd

Flg.64. O@tltt&t tt tu tw tp. d. Ac' ttrr' d' 

a. f,, .llk dr&r donrl dd rdd ot dt; c $a orr" 

B

:23t 

Holotyp.. I d. 7{V-95. Fahalabrd, 

Prntyp6: F.ielabad 6 d, 7-lV-95; Htd.r.lEd I d, 2l-X-93. 

All the speincN of thi speics hwe b@n colla&rd frco hcdsB at 3 - 4 fet 

aboec grouod while stridulatng .t nisht. 

DisringuhhiDg cb{l.teB: 

This sp@i6 3ho$ iB d* @^bla@ tlt O. crtutrl4t sp. N., br en 

e6ily be sepdatd by thc fewer numbd of filc tcdh. abdoncn enh a nattN nedian 

bowi band on do um aid senitalia with dedian lob€ alm$t trumdre at 

This speies has b4n named afr.t its stridulalory fil. wbich is ptdidcd vnh 

The qT. maErial of di5 spai* ha! ben depGit d in rn In*t Muqi 

D.p.nmcot of Agr icultu ral Enlomology. U n iveB ilv of Agr icu lture Faislabad' Pakista' 

Family Mogpplistidre 

h is ep@tt d by . sn'gL mw 3p€ks under a linglc eenu!. 

G{u Cotv.rd4 Do[v$ 

1927. G.otw.rniq Bolivd, Es, M.dr.,3:247. 

'l|'IP': Gots.dia disPar Boli!4 

A single pan ofn ie* sp.cies comPlelelv reeembles the PubliEhed d*riPion 

(chopad, 1969i Secil, 1989) of this 8.n8. 

with thc follo*ins additioir: 

Pmmton in mab slightly poduqd b&kMr& almcl r@hing to $. sfidllarorv 

vcin of .lyra (Fie. 65D).

MALEI 

-234- 

L Cotw.ndla qratuinactllt4 sp. aoa, 

(Fiss. 2E, 654-E) 

Siz. small (6.8 mm). Body broen wifi pal€ yeuowish legs. cover.d wirh dcNe 

Brcwn, .ln6t s wid€ a thc dtcrior width of Fo@tumi @iput @nExi rcrl.x 

sloping op !o die frcnral BFun (Fig. 558): rh€ larrer M'l, 3 lida a wid. ar ls! 

anrennal joint very st orSly produc€d b.twen anten@, t un@E ar apex *nh rcundcd 

arslls, v€ry 6Rly turo*€d ar middle (Fig. 654): chel6 yeuowi gena. a'rd subgeMe 

bo$ wirh a black spot belo* .yes; lac€ shon a|ld widei clyp@frontal Nturc teebly 

angulare, with two dark spors slighny above lhe suture on bod sides of the mcdiar 

furow oiiro al ro3trud. Eyc6 felbly bulging, dep brcw., Mdillary pllpi brown. *ilh 

fitsl l9o segn€ s r€llowbh. 4th sliShtly long$ da. 3rd ald 5$. th. l.rtd sliShrly 

wide.ing, obliqu.ly trunqt d ar rpcxi labial Palpi v.ry shon- AnteM brown, *i$ Palc 

Promtum 0.?3 aod 1.0 rins d loog a its ant4ior aid pccrior widtht 

€sp.ctivcly, disdrclly nm*ing in front. iL3 anterior ndgin stongly ondvc, Pdc.ior 

almosr luai8ht or very slightly poduc€d pdteriorly al middl€i dbkatm6t flal, Provided 

wirh scaucled whnish sqles, unifomly brcwn, anleriot half with 2 blmk dob along fte 

sides or the median linc (Figs. 65CD)r lat Bllob6 snall, drker than the disk, anSlqs 

.ound.d, .xtrcne i.fedor hargin v.ry ti.bly sinuat . 

Llgr prl€ yelflnh, nuch liSht r than body, pub.wni, d.r.d with abunda.t 

shirirh and gayish sls; utqior lnd m.diu fenm unifdnly @lour.dr mt no. 

ribia. pedodcd wid a t&gc, dal lyBPdun on inmr sid.; 6rti lont, bai|! ti lon8.r 

da. lhe rcxr r*o $gn.n$ iogclhcri PGErior fcmh 2.99 tim a5 lonS as eid.. bwn 

.t apcx, ribi& hther shofllr, f@bly widening losd{L apd, providcd {ith small 

d.nricles on both 3ids (Fig. 2E), adcd wi$ 3 aPrdl spun oi boh sida. th. mcdia! 

hrger hn otheni pG€rior balimrsi lhick. srcn8. @mpEsscd, 0 41 tihca $ lon8 as 

ribiae, prolided wi$ ? - 8 inl.rnal, l0 - ll externaldenticler

t.65. 

:235- 

C'.,'r.da ot tt'it ale { u A, nt dr E r'd "d 

#jiH,sl$"3nffi,'li 

i .hd.|E 9, tlodl .tt c. 

ri: 

D

-23G 

Elytla b@d. 0.93 tim6 a widc a lons, op6q!c, it! aricil n&gin ad sids 

diffBed with brown, almo$ glablous enh t@s of fift pubcs@re nelr apical @gi[, 

rigbr elyron overlapping fte lert oner mir.or obliquc (Fis. 65D)r do un finely 

oiiuSat dr lallnl f'cld blackbh, wi6 sup.rior naigin whitish. ant d wnh ! single 

blackish v.in. Wing! vdy shon. 

FEMALE: 

D.rk brcwn, d.presd. cerci yellowBh, dottcd with brown. c.nitali!dissat€d, 

ll is sinild !o $e male. but dirTeN in $e tbllowitrS chd&tets: 

Aody d2rk brwn, wilh a Eddith tinge, @vq.d wih abundant whitilh and 

gnyish s€16. Prorctum O.E3 ad 0.91 tine 6lona s iE dt rior d p6t rid widds. 

lery fecbly narcwi.g ii frcn! tloth antcrior and posierior margiN @ncNvc (Fi8 65E) 

P6rerior remo6 elongate,292 rim$ a long d wide Elvlla and wings abseir' 

Abdonc! strongty dcprecd, @recd wnh sl6, mtc stotglv on 3ide C'rci *idour 

bmwn doB, panly btol(cn. OvipGitor .t!agtt. *ilh lpi.al valv6 Eddish'blek. 

lanc@lare, snoo$, suoryly acule at aP.x. 

M€sunruots (m): ld r9 

1.75 

'.1' 

;, 

1.35

H.T,L,/H.RL 

It,F,L,/O,L 

'.]' 

-231- 

I d, 22-VU-88, Rajaipur. 

I r, 22-Vll-88, Rajaipur. 

goth $e sp@imea hrve bcen collatcd fioF a cotto! field havins c@kl and 

DinlogubbiDA ch&.ct E: 

Thi! sp@is hd a clc. r*nblsme to G dlrip.rtrir, ftom whLh it diffcs in 

6e following chtut€^: 

Premtum nuch wikr pBr.riorly thd i!5 n dir length; dirk pnid€d with 4 

blek doui l€gs unjfmly p& y.llo*ishi clyci cp.qE. ting.d wid blwnish at.p.x 

and on tidd, t. riglti on ov..lapping th. lcfi, doBl li.ld ahtost gla!|@, la&ral iEld 

bl&kish {ili bLck rciNi erci ycll*ish. dott d with bwn and thc nduenent! of 

$€rd (1989) idcnrificd t*o speim.N 6 C. alr,r'.a6, but the ahove 

diff€renB snow dEr lh.y wdc of a clos.ly rclat d ncw speics. H. ale idcnrificd a 

fenalc foo $. sam. l@lity s ^ Nw Wr.i.s 1i... Prchron biu bjdput Bis) on 

the bais of wpcrfmted micrior ribia. and frc .l rootilm, which h nor thd twie 

6 wide 6 lsr antenEl joinr. Bu ii, und.v.lopeil body chareteB sho* thar adually n 

was an immat!rc female oi some $known spe.ies. 

The name of thB sp.cics h6 b€n d.riv.d frcn ils pronotun, which n provided 

with 4 bl&k dol5 in both scxca, 

The ryF oacrial ha! b€n d.p6ited in $. lMct Meum, D.petrc of 

Asricultual Entonology, Univctsiry of AgricultlE, F.iehhad, Palisldn- 

.':.

-238- 

Fsmily Scleropterids€ 

A single spaics ha! b@n id.ntifi.d under a siigl€ gena. 

Gdu S.r.roprrer H.rn 

1E42. Scl€ropterus Haan, T€nniok Vcllundl. onb., p. 232. 

Typ.t Grrl6 (s.LtuptM) @n@.u HN 

A single @ll@cd f.n l.l&gcly ullia wilh rhc publbhcrl d*iption (Clropdd, 

l%9) of this genu depl thc suFrior rugi. or pdllriot tibi&, qhich i5 smlalld or 

deiliculacd (F'g. 66D). 

l. S.!.mpt M dcr4r8 Bnnre. v.w. 

(Fi85. 664'D) 

1E93. kleftpterus /lr.rarrr Brunncr, v. w., Ann. Mu Oenove. 33r 204. 

The only f.nale .grs wi6 the published d€scripdon (ClDPnd 1969) of lhir 

speica .xc.!t lhe following chaEcle6: 

AnEnoae wi$ t whithh ring a litd. bevoid dc middle; P.onotll mcdiat 

drination dore prcniF nar aftriot !d pdt rior mdSiBi elvca Aicnding w€l! 

b.yoid abdomen, spacd b.twen veiis fill.d wU nrcsult u'aNv.se vcinlcE oP !o $e 

middlc atd Id the apcr, lhe Bt covct€d eith ECUlatl, Pdailcl' r.slce veinleB 

{Fig. 66C): wing. very long; pccrior tibie dftd with E interdl, 16 ' U qdml 

small d.nricle. ovipdilor valves wnh l0 _ 12 snall te$ 

M.suren.nls (mn)r I 9 

B-L. H,W. P.L. A.?.W. P.P.W. eL W.L lt.F.L 

9.4 2.t5 2.75 z.fi 

H.W./A.P.W. P,L./P,P,W. E.L./P,L 

0.94 0.31 x.36 

rslamabad I 9, 8'VIl-93.

Flg.66. 

t 

-z3v 

S.|mpttu pun64rn. A, lEd [d FMo ?. dod 

rrdj B, ri6 9t C, Crh 9: D, !..fld dlt 9.

Ihblt tr 

. 

-u0- 

Ir wa @lleled fon gras undd a light source, 

This spcciB alongein thc family Scldopt ddae ha b.€n .@dcd fd lh. n6l 

& n diffe6 fton the publirhcd ddghtion of t puttldg in ^ ntnh[t ot 

ch.fui.E, it might be a w spcciq. A1 pr*nt. ir is ientatively plaed ond* rhi! 

sp.cias and its shrus will be 6ully d@id.d afiq lh! @ll@tion of iB nalc. 

Iamily Grylloaalpidae 

It ir Epr€enl€d by thrce rpeci* under ! single genu. 

Gqts erytublry t'at . 

ltoz, Gt l!o@!pa tiftille, HL!r. nrr. Crur. lB,,3: )?t, 

lEl5, C!r,/i" oken. L€hrir. Nat.,3: ,|,15, 

1913. A8nohtpa Mjohdg, Ent. Tid!kt., 34: 30. 

rr!€: GtruB lA.hrto) g,rlk alPa Lilll@s 

Thc @ll@cd sp@jme.E e cdy ully with the published desfiPrion (Cho9ard, 

1969i Ot€ and Alexander, 1983r Townend, 1983; Saeed, l9E9) ofOb ge.us. 

Key to th€ spcl6 ol G4uot4ltd l'{fmilb 

l, SFidul.bry l4th 30 - 42. M. wid.ly sp8.d * @nier of thc fdc (Fit 57B). 

_. SrduLiory t cth 50- t5,16 uddy 3P3!d r1 @nier ofthc nlc (Fis.6EB) 2 

2, Si& snau {20-5 21.0 mo)i stridulalory tcelh 50 - 52 n4l& Bum.ld.r 

_. Siblalg.e (27,0 3l,0mn)i sridulalory l*th 55 -75. patitrano s9.ad.

-ut- 

L Arylbt.lpt odtnttUt B|J'mlde. 

(8i8! 67,{8) 

l$9. A,lUotoJpa onaralir Burn ilt r, Ha.db. E.t.,2:739. 

TIE @lld.d mt iial gretly r.ui6 with thc plblirhcd det6iption (rowscd, 

1983)of o. Lr ir, exept slight kriation in m4urcmcnts of wiou tody Pdtr. 

Tow$.nd (1983).i.tcd rhat O d.r'ln h morPholosiqily itdiltiasubhrblc 

frcm C- on.tualb !d t!u! tlFy b. lyrcnymou!, Ifto, tl* lftar *@ld bc thc 

'nay 

o^ly twi.. ot GryUotalpo @nnon lo thc Afrotropiel rnd orhnol r.8ioa. 

A5 O drrri&t ds nol @ur at pr6!ft in th. ori.nt.l n8ion, ti. lp4imB 

rccordcd fren thii Fgion probably bclonS io G ,n nrdl.. 

M6!lt!Mr5 (@): !d l9 

77.1t 2.49 

5.23 0.21 

";" ";,.,. 

t.2l-1.23 t.22 0.0r

lr. 

:r42- 

pnalrr:::r-r:-r::r* 

W. 

','b 

a*bat4-l*t-tt

-243- 

F.h.labad I d, l4-VI-94i I 9, l5-VI-94i 2 d 2 9, unilbdlcd, t kei from $e 

$udenB @lldion, Depanmnt of AgdculuEl EritomobgJ, Uni!6ny of ASriclhuE. 

H.bit.t: 

h wa @Uet d fon dc mobt sil along th. b.nk of w.t r chels and frcn 

ClDpad (1969) and TowNeod (1961) menriorcd that A dti@ does rct 

l:M. in tuia and i is app@ntly r.pl!.d by G, o,i.tualit. 

1E39. 

1906. 

tv25. 

2 GryUotalPo ntuuta Bln w{ 

(Fi8s. 68A8) 

GryUdalpd nhuta BuincisEr, Ha.db. Ent,,21 740. 

Curtilla ninaa Kirby, synotr Cat. Onh,.21 6 

Gtllotalp4 ni4ma vd. Cti.re Chopdd. Boll. MN Hist ml' Puir' P.442' 

Th. tm slle&d nals fmm $ir region gdrlv @nfmd !o thc plbtish'd 

d€scription (Towieid, 1983J of C d.biUt, b diffcr in the nunb.r of sridulabrv 

c.rh, whicft arc 50 - 52 lnd lB widclyiw€d eit r or thc fiL (Fig 688)' 

Towns.nd (19E3) ert d th.t c. d.r/{ E @nfus.d *ith o' '!i&'4 burdifleR 

chiei'ly in thc fcwcr Dumb.r of stidularory 1eerh (hcrn 31 5)' which e 63 in the 

holoryF of rh. laner. Hc lunhd mentioncd lhat C nnlra is 6nmon in tic Ori'nt'l 

rcgaon, but dG mt (Er in Aftid, In vi* of the above Enakr, lhc sp.cimN 

colle|.d fron rhis rcgio. arc idcititicd d C tirlr4 fd th.v core cl*r lo the 

M6uEMls (m): 2 d 

B.L

n!,44. 

-* 

qOyt****t*oo*** 

alrtL|'fr.

245- 

2.t! 

t.:26 

na 

Fdsalabld I d, 2GYI-94| I d, unlab.lled, lrLen aom th. ttud.lls @ue.tion, 

Iteplnrn nr of AgricuhEal E romoloty, Univ€rstty ofAgrtllirc, F.j$hb!d, Pakji.D 

MALE: 

0,t3 

o.ul 

2.O 

0.01 

ll h|. b..tr @lLcsd fiom vcty sa loil in a nltrrv. 

mi! 3pEis b6 ban @ordcd for thc fiBt linc fdn lhlilt !. 

3. ArflottlP. PcA$M sp.t$. 

(Fi8r. 24, 694-D 

Si& LrSe e?.0 - 31.0 nn). Eody coloun6o &tl br*! wilt l|g!t rqH![ 

rirs., nrcly U&k!h+@n, d.iicly Puh*ctnr 

Smll. 0.6l - 0.68 tiB s wid. r! prelotm, stronglv onicd, bl&kish+rcM 

Mouthpins with labruB brown, clyFu ycllowi lMiuarv FlPi with fi$l 2 toi s 3E ll' 

y.llNirh, 3rd b|oM, llr86t, widdiog lo$!flb 4.r, &n.!d 5th ulcqEt 6lh v.rt 

sn.ll, dMr rcld.d; lzbitl FlPi !!dl. Eyd p@rrding, bl*l(' m.tlv reuDd.d. Oelli

-216- 

trc. laBe , o!!l, y.lld (rely cd in bl&k spc.im.$). Anten@ btdt, with !€8mnis 

ro!nd.d, havily puhsc.nt, 

Prerctln !ary., aln6t oEl, @.owing lticriorly, 3id4 ra.d.d. atlicrior @8in 

depty concavc, pcr.rior r.Suldly @und.di dbk coN.x, Elv.ty, *ith a chafut rirlic 

pattcm in niddl. (Fig. 694); lat.nl lot ! fc.bly y.Uowbh dong i.f.rid Mgin, 

Lls3 d.ely pub@nti thc a.tlrior sPeializ.d fd dirgiDr (Frr, 2A), blwn, 

rinScd with tlddirhi mildle lcg! sFall, y.ltowth: p6arior fcmo6 d.tt r .long 3upcriot 

hall rhc rclt liShar; po6t rid tibi& shon, rrmd with 4 sPiGr o! idc.n l naryit, 

Tirl P'rB ? (4 icr, 3 orr) (FiS. 59D), 6{Foi ..nd! ldU6t, strDr! rdv ildt 

Eltta m@h rlonq th.n abdomni sridul.lory aE oblo.t, ! 96 - 2 25 tima 

&! long ai {idq dia€or.l vein sligldy cuN.d Fe b.ei r.diu divid.d dbally i , & 

aid & (Fit. 698), trltlEe Einlct! 16 pmnimnt, fqdiig.loiUra elk h apiel 

f'cld: 3nidularory filc with 55 - ?5 l4$, lcss eid.ly 3P4.d n diddlc (Fi8 69D; lll.til 

fEld wiln sc bcuiry 7 - E b.rnch6, .ll Pointing lovrrds wi[t fiP. lst nldnlr barh.d, 

t aEv.E vcinl.b P@nt along qlr.tu itfcrior ffi8in. wing! neh longer th2n 

abdomcn, 1.95 - 2.25 dme. as long.s .lyt6 

Robu$, brdn .bov.r y.llowish bcMO. ccrci bns. brown, plb.sent with loiS 

Mir. G.nihlia shon, wilh peud.PiPhall6 r.fowly @diforn, bti.8 Mw b|4h6: 

vcntBr pr@rs rarh.r 3hon ad &urc (Fig. 69F). 

FEMALE: 

It is 3imilu to m.le erc.pl th. followi!8 chdollE. 

Elyta qith doMl Rcld having r ncr disb Eia, fc.bly cwcd al b.e€3, .lndt 

Egllady spNccd it .Pic.l hatfi usvqle lcinlcts Nncd3, fdDi!8 ld8. @Us in basal 

half, llongal. ltd wow @ll! in .pi6l h.lt ndiu lom.tiG divii.d dilhllv into R' 

a d R, (Fis. 69C). Post rior libia. dn.d.bovc wnh 3 _ 4 ryiB 

on irE nal d.rgin.

Fa. 69, tub. t,r'dttutt F. d. A',.n.4 'k l+t 

& -drd ;, d.;l .!d Lld'l .ad' c, t|i u tr D' !4qrq 

tlbb J! f,,.trlddd.ly fih Jr F. pLllc o4|.r d. &d

M.GltEot3 (Dn)r 5d 59 

ll.LLr,It.F.L 

27.G31.0 

1t,2.r2.3 

Lta-l.32 

-24V 

2.29 0.13 

3,t0 042 

26.G3t,3 

7,L4.2 

r.t6.t.x 

l,l3 O.,l{ 

2.12 0.19 

o?l o0l 

Hobr!?e I d, t&lv-9, Mllra. 

Allolyper I 9, lElV-9, Mutlan. 

PattE6: B.[.sdpu I 6, 2GVl-EEi r2rnrt | 9, l6vt-Eq Mdrrn, !?-V- 

9i Murcc I 9,21X-88r Srrgodha I d,s-x-EEi Siatkor I !,2E-vtrr-8Ei l0 d 8 9. 

u.labelLd, irtd ftdl rhe stud.na' co .clion, D.p.tuEnt of ASriqjtu rt Enionotogy, 

Univcsity of Agiq urc, F.isrlabad, Pakhran, 

Thc ape.i* ha! be€n conbonly colt@t d on lighB and rarcly &om the banlc of 

w.tcr chanmlr 4d eals.

ttdlsut'bItrg.Drres: 

-24v 

It kcy! our ir Tow cnd (t983) eirh G rrrrd4,bul ditren froE it! pubtiNh.d 

.leeription in th! NEbq of srilul*,ry fil. Edh, stid u. td *i&ry-rF..d i! o"e 

anrlr. h is dlo wry clse b c. dn.ara, but difiqs in mr bavinS the ch!rcicrisdc 

mlc S.nitrlir ai in rh. Lc.r !p@as ad ale in rhc nunbd of snidutatory fde EcO. 

sud (1969) cfrFtllly id.nrifi.d 3 mat6 rd I f.n L a! c. foto. t n 4 

rentJ6 a A. 1lJ.1. Th. a!!lr!i! of th.i. trelc gcnindi. ald numbq of f,te ccth 

during the pr.l€ii r1udi6 r.v.ded d[r rhcy &tudty b.tong.d ro a n w sp.ci.s. 

Tle nan€ of this spc.is ha! bdn dcriv€d from prkilcan, rhe mme of rhir 

Th. typ. EErid n8 be. d.po.icd in rts IB.cr Mru. D@.ncn! of 

Acriclltlral Edto@logy, UniEsity of ASicul&r., Faisajrbsd, pstbtan.

SUMMARY

CIIAPTER V 

SI,]MMARY 

Thc adult crict b wcrc @ll4l.d du.ing the ytus 1991 lo 195 fron 46 

bcalnis of$e four clirstic reSjotre of Pakisljin. The @ll@tioN *e@ suppLmcnicd 

with sEiftN of the ldiou3 eork.* 3 wll 6 dE snddt'' ouetion of dE 

D€perrnenr of Agrioltuil Enbmlogt, Uni*6ity ol Agri.ulbe, Falalabad, Pa*BBn 

Th. sleincN wc ideiftlcd inb 6E tpe.b ad 3ub6p@i6 b€longinS to 2E 

8cn.a ir 8 fanilis unds th. sup.rhFily Crllloida our of rh* o.. gcnu 

PholliLbot6, ^nd 31 s|6ies, riz,, TutabgryUls kaLw6is, T. buUahi' T. 

ptodiM, coto8ryhd oviLryL!, Tele%ryU!$ li8nd6, T to66, T. Pt@.tut 

T, lnneuidd, PlotreryUlA biconptrc, malblobosu adoitiualit, P tuh4\r.Bi' 

A.heta thotrcica, Plcb.iosrtUut t tit.Eubnt, lsoblefu u.aqpLt' L. 

potohtrcNi!, L. dolichop.nnis, L. "Edior.Pluka, L Ms4low, L .ot$6u' L 

dMl6, L. pi&46, L. hmihMalat, V.l^tifcro6 4lveol6, v . Ilni,ot6 

V. dn tuN. V. @tuirt6, V, .livd6, Mo.li@gryllu bo@14, M. cotdtttt 

M, ttlodispaB, AM$B kanalu, M.A@,Pha n@ Mla M. @nul'1t', 

O@th&J crcnortg, O. t!4nupo6u' Cntw.&ia qudn"verlata td GoUototPa 

paristaaa, e w t4*C@.'flr..lld.ypcLhaLof Loxoblew hM tu haalh 

ben d@ibed for rhe iBt tide fron its lype l@lilv 

lr addnion, the f.nily S.l.toptcttnte, 8 E n n (C4lloeryl6 T.Logt tIlB,

-251- 

Plcb.iotryllB, TananSryUut, E tcytr6, AMutsut, Anatipla, S.ktoptewt, tl 

sp.cies, such as G,tlaptit Ubetcen, T.boguUB lontip.Mit, Ptatsrytu 

Mlaw.phal6, TanarcEryllB hnatu, Iôblzw ho@i, V.tttifidot$ 

6p.66, v. Ialkt, Ahuipha toryip.nni!, Eu.t !tt6 condnM, ScLtoptet$ 

padat6. GryUotalta ninut4 a afia|'ol Ta'riNkicltg pon. oflrr &c r@rd.d 

tbr the fi6t tim. iroF Pakilhn. 

The cotr4t idcntifrcadon of l7 sp€cks. which erc prcvioBly niridenrifi.d 

by Sa€cd (r9E9). is siv.n. A singl. holo{r?e n.l. dcrrib.d as I@U.M jindti 

is synonynized rifh L. hanzlif.L me ideaircîon or Cophos'lqut btuiry ^rA 

trE n* statc of P@Jttonabi8 tui@1pu6is d sivcn by dE alove wort r e 

doubtful, a $.y de bo.d on v€ty ihmatue spc.imeN, which l&t rh. inpoll! 

Th. charccrs of dE fdily,goup h.vc mt be. ircluded, for they @ 6ily 

a.essible in lit rarur., b kcys for both lhe higher and lower t ta have bcc. pr€p.r.d 

in oder lo facililale thc future idlndfiqtions. The deviarioN frcm th€ publish€d 

dscriptioa and rh. addnioEl cbd.E6, il any, of all rh€ g.I@ $d speiB e 

included- The ncw g€nera, species atrd allotypB hale bei d$sibed in delail, The 

n*urcnenls of variou aromniqlly iopona body pa.rs .lorawith lheir hngc, 

meansi sllnddd dcviatiom and ratiq havc been wo*ed out, Thc iuusrr.rioB ofdiffcEnl 

body pats alon8wilh lhe g. lalia ard dhributional nrps ol aU ric sp.ci6 de lls 

giv.n. .Ihc lalitu&, loogitud., mu.l mean bnperaoE and annualav€68. prccipiution 

ror almct all th. oll4tion l@liri* have beo aDoended-

CHECK LIST

CHAPTER 1I 

CHECK LIST OF GRYLIOIDEA OF PAKISTAN 

. 

Fs$[Y GrYIU&e 

Subf.n Y GrYliDre 

C6E Trturd.tdlr. G.r.Ltov 

L l ,ot4roi6 (Lrc[e!8.ir) 

Galt Eltta'r&t Mrh 

2. A dior (Cl'q6rd) 

Go6 tt ltott l&, Tdu6rt 

t. T. tdtiMtw (t ifct Alghlr) 

1. r. pa*itt,,ut (Chourid Ahtr'ld) 

5. T. ItlMit q,N, 

6. T. tullahi .P. w. 

7, T. pmdi.6 3p. nw 

C€nqs C4lbrryrr,rt Sjocledl 

E. C ovriL'AA sp. nov, 

Gdu! Cttlrt rit Cbop.rd 

9, c. p.rdca CIFf6rd 

c.r'6 C'l&, u|ltlas 

lo. G- biwtldtt D.c*t 

G.nG td.olt rr$ Clop.d 

ll. r lied 3P. w. 

12. r b44att t ($!$u|l) 

13. r )ea4t ry. mv. 

14. T. prmns sp.w. 

15. T. hrsr us q.N

16, 

t7. 

t8, 

31. 

38. 

39. 

40. 

42_ 

43. 

:u3- 

Ga|r3 flrrtlrter Choprtd 

it ttwdetur (sctaitt ) 

G.nu ftulolrrorlr sq. nor. 

ad.ntibialis sp. t$\. 

A. doMtticw (Linwa) 

G.na Acl.r. rrbri.iE 

Ge.a t.r.r4rrl&r Rfl.bll 

P, Bieautuit sp. w. 

C.nur O.rrbd.r Sr6W 

A. suWUcM {wat} O 

Ge.us lolt4Dlrr4s T&ilBkll 

L. pddan6is q. oDv. 

L. doli.hopntis sp. ia!. 

L. n dt@c$attt sp. nov, 

L. h.nihwht q.w. 

V. r.tiQ@t6 W. ^q. 

Y. /rrh (c!oprd) 

Gou! IerL@ S.|Ec 

G.!6 Y.L'f.rom R.lddl

-25+ 

G€nu ttod.or.r&r Choprd 

44. M. ,lrrM (Sau$urc) 

45. nt ri8rt a (Walt r) 

a6. M. .ht@ lcf6p.rd) 

17. M. b@tlB sp, w, 

44. M. @n lts V. w. 

49. ll. ttldispdis sp. ft|t. 

Subfcmily Nemobiinae 

G.aa n@4.nobi!,3 J.@h6or el Bi.a.H 

50, P. @r@/r. (walt r) 

GenE Dlar.ror!6 llcke4 

51. D. .rilli (Bolivsr) 

s2. D. f@ipa lwalkct) 

FaEIt TrtgoDiliidee 

Cinus ]{r!lr4!, E|t,!!F @ Wdtdritl 

53. A. kanatg q. N. G€nu! lhigorlt D Rsbu 

54. T, ci.irdzL.kta Pablr 

55. T. c. hMb.niarun lsureveJ 

G.nu i@hl4 s.|sn 

56. 'r. basip@n!' $wnb) 

Fdaily Ph.lngoFnd{€ 

C.!0i lr.r.aiot rd wdr.r 

57, M. @lotata +.w. 

58, M. anttTa q. w.

59. 

F.mily beo@idae 

sublss0y Eaeotserinre 

Gsxu Xdoltt&|, Iotrt.r 

X t r,lrwBn (Wa|r.i) 

Subfam Y m@irtinte 

Galt &rct t Gcttu{.a L 

60. R.o,tci'ru (H!rn) 

61. o. 

'62, O. 

63. O. 

65. 

66. 

a. 

68. 

e- $dtitulld. 

F.sily Oe.r| Didr€ 

G.dt6 o..-crt!, Sdt{r. 

F.Dily MotoPlhdd.e 

G.!u C,o,!td, Eolitrr 

q. w. 

F.nily Sc-l€mDf.rfdr€ 

G.r!s tct roldtt s|! 

J. ,lrrddB Bruds v W 

F.nilY GrY Dtrhitlre 

G. E @/.be$. rdD! 

c. ottatalis Bvafirts

LITERATURE 

CITED

CTIAPTER VII 

LTTERATURE CITED 

| . Abdull.!, M., 1955. Cryttitt . of Lydtpui. M.Sc. Thcsir, Utriv. pud.b, r..toG, 

2. AluEd, K.S.D,, 1951. Ctif2ric 'rgidB ofw6r p!ti!rb. pE. lrd, p.r. Sci. 

Cotrf-, 101,135. 

3- AhMd. T. atrd C. Ufr.h , rg4g.a'tet-tuznd cict r .AryAtb doMti% L., 

a a p*r of clops. prc. l3r. pat . Sci. Cont, pr. 3, 34-35. 

4. Ald.ldcr, R.D., t991. A di.r of 6. gcn! Crl!(r (OdopLn: Cryuib.) 

pirh a icw !p€t* fion KoM. crer t*B Eni, Z(2): 7}Et. 

5. Anonrmu!, l9l-73. Co.Eot of qttcB h Der. ch.ri Kh.r iftri.r C.ick r 

Corrol Schc0c, 4 pp. 

6. Anw$, M.P., c.H. MuBi, T. HUgh a.d M.r. Staiwi, l9E?. IE.rFr, 

a$ociar.d wirh posto ciop at Taidoj&n. proc jth. pat. Cong. Zool.. 

l49 l5t.

-251- 

t. Barerje. S.N, and B.K. ChaGrje, 1955, Sudics on C'llk alpo 4ric@ 

Palisr de B.auvois, a iew majo! pest or the Potalo crcp (JoLr@ 

rl,.Dr@ Linn.) and irs @nllol. Ind. J. En!., 17:219-229. 

8. Bharg.va, R-N., r98r. A rcw 3Peci6 of s.letupt M (Han) (tneeb: 

Grylloide) trcm Nonh'WBt India- C@bio6. lodhpttr. E(2): 57'59. 

9. Bhaqava. R.N.. 19E2. A iew spccies ot Luobtenn6 Sauseurc (OnhoPl€n: 

Grylllta.) fton Nodn-welt India. Oe.Dic6, N.w Re? 9(l):4&49 

10. Bhownik, H.K., 1967- Five mw 3p4is of senus S.CAltP.ld Sausurc 

(tNet : Odopllh) fioF tndia Prc. @1. s€ qlcuna 20: 123-127 

11, Bhownik, H.K,, 1968. Six rcw sp€cies and one n'w genus of Grvllidae frcm 

lndi.. Pr@. Ind Sci- cong. As , Bcuas pt B 49l_49 

12. thowmik. H-K., l9?O Tne glvllid fauia (OnhopteBr In$c1a) of thc gaat 

Nicob.r lsldd, lndia J zool S@. lndia 22:69_85 

13. Bhovmik, H K., l9?1. Two new spei* ofclvlloide 

Oieit. IN.,5: 503-506- 

(Orthopoc'a) fron India' 

14. BhoqFik. H.K., l9?5 Studi6 on Indian crickeb (OlthoptcB; Ircda) pt lll' 

J. Bonb. Ml. H!r' S@. ?2(2):368-382 

15. BhowFik, H K , 19?6. On thc grvllid rauna (Grvlloida: orthopEB) of ibe 

disricls ofNodh Bedgal with ddfiiptions ol3 req spe'i6 lnd Mls Brll ' 

ll(l): 42 'lE.

t6. 

l?, 

18. 

t9, 

20. 

2t, 

22. 

24, 

_258- 

Chopdd, 1., l9l. R?on on tll. Onhoprerr of Mcsopobnia ad p.Bi2, 

elle.ted by Mds^. p.A. Buron, ad W.E. Evas, B.Sc. Dicryofl.ra and 

Eisifera. J, 8onb. nal. Hi!r. S@.,2i.751192. 

Chopad, 1., 194- Otr $de @Eoicdow Orrhopt B and D.nnaplera fon 

Assan and BurM, R@. Ind. Mus,, 26: 8l-92, 

Chop$d, 1., lr5_ Thc Cry id.e of Ceylon i, rhc Brirish Mlseun 6[e.rion. 

Ann. Mag. iat. Hbt., 9(t5):505-535. 

Cha&d, 1., t928. RcvEion of d. Indirn Cryrlid&. Rec. Ind. Mu!., .t0: rJ6. 

Chopard, 1., 1936. The Tridadylid& aq, 

20: Y87. 

Cryllide ofCeyton. C.ytor J. s.i., 

Chopard, L.. 1959, Cryttides d kan. Slulrgart. B.ir. z. naoR, num,, 24: l-5, 

Chopdd, L., 1960. Conrjburiotr s' t.erudc dc h 

cryllidcs. Eos, Madr., 36: 389-40t, 

F UE d Alghrijur l5i 

Chop.rd, L, 1951. t4 divjdioB du ecrc O.}lk, bas.6 sur lcrld. 

I app.Eil copuht q (onnopr.E crylli&r). Es, Madr., 37: 267-26?. 

Cnop6rd, 1., 1962. hlop(.G cryIjd.. .1 Cry eridida. d. l,An8ot . pubt. 

cult. Conpad, Dim. Angota, j6: l3{9, 

Choprrd, 1., 1963. Nolls ir t. faurc d6 onhopcmids d,tsr.eler dc qudqB 

r.gDtu tinirophes (Blrrioda, crylluidoida, crylodo). Bun. R6. 

Counc. ls@I, lt B/4. D. 169.

-259- 

26. Chopdd, L.,196?. Cryllid6-Fam. Cryllide. In Beiei, M. (Ed-), Orthopl!rcrun 

C.trlogut, l0: l-21L 

27. Chopdd, L-, 1961. CryUid.s-F.d. Cryllida., Oe6rthida., Gryllotalpid.e. lo 

B.i.r, M. (Ed.), orrhopt en Cal.losu, 12:213-50. 

26. Chaaid, L., 1969. T1E tuun or Indh art adjent @ ri6. O.thoPca. 

Vol. 2, Crylloida, Zool. Sufl. India, Calcutta,42l pp, 

29. cody, D.H.L., 1946. ,lrLtru (Hcniptn: Lrla.id.c) in sro.€d Glo{rd'nuB. 

Bull. eDt. Res., 37(4): 609-61?. 

30. Fe@ndo, w., 195?r. N.w rp.ci4 of lngts frcm Ccylon (l). C.ylon J. Sci. 

(Eiol. Sc.), l: ?-18. 

ll. Fer6ndo, w., 1957b. New sp.ci6 ol InsEt! frcn C.ylon (2). C.rlon J. Sci. 

(Biol. Sc.), l:73-?6. 

32. FerMndo, W.,195?c, Contributios toa knowledge of tie lMa ofCeylon,2, 

E tltotkw taftltueotui sp. nov. Ceylon J. Sci., (B) 25: 201-202. 

33. chosh, C.C., 1912. Thc big br*n ctickcr (gru.lqtrtryt @h.nn6 stoll). 

M.n. Depr Agric- lodir.4r 16l-1E2. 

34. choufi, A.S.K. and H. Ahmad, 1959. A rtudy of th. crick r! (Grylloidea) of 

Pttili'n. Prcc. P.r. Sci. Cont, ll(3):49. 

35, Ghouri, A.S.K..nd J.E. McF.rla*, 195?a. R€pr.ductive islation in 6. hous 

crhkct (frlropraa: G.yllidr.). Psy.h., 64: 3G36.

-2& 

36. Ghoui. A.S.K. and J.E. McFarlane, 1957b. Sp€htion in th. hour! ciict l 

Achao dotuttitu (L.) (O hop|co: Gtyuid&). Pre. Pak. Scj. cnni, 

7(3):51-52. 

37. Ghouri, A.S.K. and J.E. McFarlane, 1957c. Exrra{rdinary d€velopmeft of win$ 

in cryLbds nqik"6 (\Ndk.) (Crylida.). Ploc. Pak. S.i. Cont, ?(3): 52. 

3E. Gnouri, A.S,K, and ,.E. McF!rl.n., 195&. ObscdrioB otr thc d.v.lo?n€nr of 

crickcrs. canad. Enl., 90: 158.165. 

39. Chouri, A.S.K. and.l.E. McF&larE, l95Eb. Occutrc@ of a @rop€rou fom 

of Grr|odcs siei|ata (walt€.) (oilopt ri: Crylli,r.) in labrarory 

cultue. Criad. J. z@1., 36: E37-838. 

,r'0. Gorotlrd, A-V., 1978- C.iclcE of the gc'@ T.trmgtu!.t T.tb. atl.t 

Mo.licogrylta Chq,lo.Jl'.4Jtr;n: C.yllid&) in niddL A!'a. Enr. Obad., 

57(l):9.105. 

41. Corckbd, 4.V,, l9El- A r.vicw ot fi*ea of $e Ne4obii@ 3ubfanily 

(Onho9c6: Gryllid&) of d. U.S.S.R. tuud. v6rr Z@1., 0(2): 2l-26. 

42. Cookhov, A.V., 1983. Contiibution to the kmwlcdge of dickeb of thc Fibe 

cryui.i (Olthopleh: cryllidre). Enr. ohoz., 62(2): 314-330. 

43. Kevm, D.K.MCE., 1980. The OnlDpr.roid iMcE of the B.0nudas. Men. 

Lymar.nt. Mu. R8. L$., 8r l-182. 

44. r\hn, H., f 949, Bicnodi6 of bldr-naded dict r e ryUtIB doMtt B I',m- 

in Sind, Pc, lst Pa,l. Sci. Coni, pt- 3, p. 34.

.26t- 

45. Kirby, w.F., 1905. A syrcnymic eblosu€ of Ofthoprere, Vol.Il, Ofthopleh- 

sal6ro.ia. pr | (Achelida. d PhasoNid&), Lrnd_, 562 pp. 

46. Latii A. and M.A. Asghar, 1957. Two rew spei.s ofcryloi&a. pre. 8th p.k. 

Sci. Conf., p. 69. 

47. Marari, S. ad N. Otaru, | 963. Phoropdiodic @nuol of grsd and qing,fdn 

i. N.nobiB ,s.or.6n ShiEli (O hopE6: Crytlidre). Konryu. 

3l: 1C26. 

4E. Matlad, S.B. .nd J-E- McFdLrc, 1968- Two .ff6 of phobp.iod otr wing- 

dev.lopded in Cry4od.r ri,gittdrB (Walk.). Caud. J. Zot.,46:5?-60. 

49, Ma$edy. E.L., 1981, ConratinS fedlnB habii! of pest nole crickcr sp4ies. J. 

Ea. E^1., 74. 444445. 

50. Matsuua, H,, H. Oda a.d H. Ishiati, 1985, Darnage 10 chims Yan by de 

afii.a. moh cricke! Crylku.lpa $ticata Palhot d. Bauvois, and ils 

6t|rol by chemi@h. Jap. L Appl. Enl' & zool.,29(l): 3@. 

51. McFarlane, J.E,, l%4. Feto* lffe.tiiS growrh and wing-polymorphhd id 

GltIod.s si8iud 1t (Walk.)r Dietary potcin lcvel dd a possibt..ffar of 

pho(lpcriod. Camd. J- Z.ol., 42: 76?-?71. 

52. Otte, D. and R,D, Al€r..d.r, 1983. The Ausrdi& cdct B (Onboplla: 

Cryllidae), Acad. nat. Sci. Phil. Mono|6r.,22,417 W. 

53. Ofte. D. aod W, Cade, l98la. Aiiid c.ickets (C.yllide). l. T.l.oSrtUB of 

6t m and eu$eo Afrio, P.@. Acad. ut, S.i. Phil., 135: 102-12?.

54, 

55. 

56. 

51. 

58. 

59_ 

60, 

61. 

one, D. ard w. cade, 1963b. Aiii@ qict.rs (Gryllida.). 3. Otr rh. Aftid 

sc@ia of v.ta,if.it8 Rrndcll (Grylli@: ModicosryllinD. PE. A@d. 

El. Sci. Phil., 135:241-253. 

ore, D..nd w. Cade, 1984a. Afii@n crickcr! (Cryllid&).4. Thc Scnus 

Pt ts?46 frcn e'llri &rd $uthetu Af.ic. (GryUi@: GryuinD. Pc. 

A.!d. ut. S.i. Phil.. 136:45-66. 

orte, D. and w. cade, r9E4b. Airic.n cdcke$ {Gryltide). 5. E !r aid souti 

Afrimt 3pcci6 of Modl@tryL\a ^!t ermt t.t^ra seE6 (Crylli@: 

Modicogryllitri). ha. A.!d. ot. sci- Phil., l36i 67-9. 

Otle, D. and W. Cade, 1984.. Afii@n crickct! (Gryllidae). 6. nc gcnB Cry{6 

and sohe relaEd geBa (Cryllin& Gryuini). Pr@. Aod. nrt. Sci. Phil.. 

136:96-122. 

Ott , D., 1987a. Aiii6 cickcr! (Gryllidr.). 9. N.* gened and !pei6 of 

B@hytrupift dd Gryllinac. P@. Acad. dt. S.i. Phil.. l39r 315-374. 

Otle, D. , | 987b, Norerctaruhl iait,shakB sysr.nuric dog, R€rly ro Dr. Kctu. 

Metakptc., 9(D: lGlE. 

Pajni, H.R. and M. Kalra, l9EE. Funher @nncnF on the Indr$ spahs of rhc 

E .oe TuMeryU8 Tdbi6kii qirh d*iiption of 3 nc* lpeies 

(oiiAr6: Crrllid@: Orylli@). Ra. Bult. (sci.). Pb. Univ_. 

39(3-4): 181-185. 

Quayymm, M.A., M. Raman and K. Ali, 1987. A ph€Etic appoach lo lbe 

bromny of Gryllid@ of Faisalabad ar Scneric lev.l. Pr@. 5rh. P.l. concr. 

Z&1.,2n-7U.

-263- 

62- R.nd.ll, R.L.. 1964. Tt @l. Scnilrlia in CryUi@ (OrthoFcF: Gryllid!.) and 

{ nibal r.virion. Ced, Ent..96(12): 1565-1607. 

6r. sdd, A., 19E9. crickers (crylloid@: onhorcd) or the Punjab. M.sc.Th6b, 

Univ- Agric.. Faislabad. Paldrl'n. 

64. s.ndrdag@, T,R., 1954. check lisl of the Ttid&tylid& dd Ctyllidre (rnecla: 

OnnopEn) ofccylon wnh.ccords ofdi$ribution r. Bonb nfl. Hist. s@.. 

52.544-562, 

65. 'Iandon, s.K. and M.s. shishodi., 192. Notg on tlE @llalion ofGrrlloida 

(OdhoPEn) fron NEFA, India. Odeit. lN 6(3):281292. 

66. Trndon, s.K. dd M s. shishodia lg?4. D€sription of teo ** sp@ias of 

V.lqifcattu (oflttoPt a: Crvlloid.a) frdm l ia O'icn lB" 

8(3).299-302. 

6?- Tindalc, N.8., tg2S Ausralian nol.{.iclcts of tlE fanilv Crytlohlpid& 

(Ortnoplla) Rd- s- Autnl Mus. 4: !-42' 

68. Toru, R.8., 193. Morc wingcd f.nalq of te c'ickel erllod's turytica8 

{walk.). S. Arr.l zel ,28(2)r l22l2a. 

69. TowBend, 8. C., 1980 A urononic stldv of two sinild sPe'ies ot Teleosrylus 

(OnnopE a: Gryllida.) J. nat. HisI., 14: 153162' 

70. TowMnd, B.C., 1983. A rcvi.* ol $e Afrotropical note+rick'E (OrdDp6': 

cryllolalpid&). Bull. Brn Mus. nar. Hist (Eit ) 46(2)1 175-203

-261- 

71. V@tt, M., 196. Thc lnknowi f.mal6 of cdct4! (O.thorl!a: CryUid.€) 

12. 

77. 

75, 

76. 

from Norlh-Frst India wi$ mtca on th.i. diBt ibution, Enionon-. 

3/J). n2-276. 

V.sth, M.. A.R. tdid, S. Bbv$ lnd A.K. C6h, lyrs. Thllc rw rDccies 

of Gryuidi. (II!.ch: Odhopcn). Orierr. la., 9: 221-22E. 

Vick.ry, V.R,, 193. NotB on n Dr.r@rl8 ald a n v genu of $. ribc 

Pa@Enooiini {On!acn: Crylli.L.: Nambii@). C.nrd. E , 

105(3):419424. 

Walkd. F., 1869. C{.log!. of thc lp6imB of Ddmat0.6{.ftaroria dd 

3uppl.roc !o rlc 8lara.ir. in ri. @lletion of th. B.iisl Mueon. Vol. l. 

Cryllde, Bhndi&, L@utide. Brit. Mus. hbl., Lrnd., ll? pp. 

walk.r, T.J.. 1967. Rdision of th€ O.e.thjn& (Gryllid@: Ont@t d of 

AMaa, surh of rh. utriEd snils. AD. cnr. soc. AE., 50(/t): 78/f95. 

YutrB. M., M. You.uf rd c. ,ildi, 19E0. ltBcct rrd spklq 6ic pcaa ofcoson 

in P.kiltrn. Fi@l Repon, P.L. 480 Projcct, Univ. Agric., F*alab.d,

Distributional 

Maps

N 

roh.r* s|4! r. 

'o 

r), 16,2r 

r ,n.'!ll]& e. ciqd) 

,f|@q'u@(|^lab6^ 

I T Frtfu (GbMd ^ldI 

Ianb'rykonr.rPa'e' 

3'ad|@bPbw.td||d 

9aqMbneal.fuD.%| 

|afd..'afu|i.fuqd' 

r! . r4rifl& Frslr) 

,265- 

Didrlbutiotr ol Crylim tPiordrltrd, t@&Brrt&r, 

C.UoE.rUut. e.rllortb. eryUtt, t.t@z,rUut, 

PhLtbbbow, kh.tr dtu aryuo.t ti 

W!.r 

') 

12. r&.t,tts Fa4r 4 u. 

tr. &)1116 r.boarroar (s.dilr!) 

{. 1.r& tulis (Lbo4) 

4. 6Jtu 't8t4 AteF'

hl|,d'4'5'67'9'|,'|4' 

ToLTlrsq!6'''Io'II'|2'b'l'} 

-)66- 

M.p2D. G.rll,/Ie tTtt uoattu!!, 

v.tadfiatttu dn 

I r44u3& uud (nurEj 

.'L'pdduek4M' 

I + ddd,P$is 'e.nv 

. L r@drLg I. w. 

t3. v.b,iteu tu;L, 4. w 

)1 vtblilttury dFu (wttql 

xMdia'414d|'ru|s'v'!rc' 

6. M DtlCj'r'tt re.8t.

N 

-261- 

DErrtbudoD of N.oobiilre i! 

hcw@da adot (,!bn 

Di,n6dh4 tdcirs N$r..n

N 

-268- 

Dlrtrlbuttd oa TrtSontdt.rr. b p*bt !, 

''1M4Bk'n.,gb'M' 

2d'o1ulhd.bd.buqn6\ 

r. r c h$.dd,ree) 

., a@pr. rdat.tu (edio!)

N 

-269- 

Mlp 5. Distlburio! or Pb.hlsopdd&, Mosodbd.r.G ud 

sdq!!...id.. ln P.&ru 

|'.|.ibDr.n'd'fu6.M' 

z. u{,e,'' tud44 b.;N 

1..*'.b.e'&.'fua'd

N 

-270- 

M.D 6. Dtutdbdbn or Eidpaqld& ln P.$r.a 

l. lr.16&. ,iur8 ($/ {) 

2. Prc(r& dfu 0|I!)

N 

-271- 

Mrp ?. Irdrlhrtlon of O.crrtddr. lr Prl|nu. 

2'oc||'|d'bE'a- 

''o'.e,.,ga'a'

N 

Mrp E. Dr*dbutlo! of cry|roa.tddr. h rrUrt r. 

' 

'.c''dfu4'M' 

q'dd.t, dde elde

APPENDD(

Nj;" 

LJntdq lddrtd4 dlltud., .drc.rE .rd tn a.ll oa l!. 

oU.rtio le|nG d Prl&rr 

214UN 

63-35E 

xl5 

1216 

9{N:M4qNh''*tuful*fl 

qwl

IN THE NAME OF ALLAH THE COMPASSIONATE, THE MERCIFUL

You also want an ePaper? Increase the reach of your titles

Delete template?

Save as template?